Alfred Russel Wallace : Alfred Wallace : A. R. Wallace
Russel Wallace : Alfred Russell Wallace (sic)
On the Pieridæ of the Indian
Australian Regions (S127: 1867)
Editor Charles H. Smith's Note: Read at the Entomological Society of London meeting of 18
February 1867, and printed later that year in the Society's Transactions series. My thanks to
John van Wyhe for allowing me access to the part of the transcription of this work at his
"Wallace Online" website that I had not already transcribed (this appears here with additional
text corrections). Also, thanks to Roger Beckman and Margaret Janz of Indiana University for
providing the scans of the four plates. Original pagination indicated within double brackets. To
link directly to this page, connect with: http://people.wku.edu/charles.smith/wallace/S127.htm
[Read 18th February, 1867.]
[[p. 301]] The Pieridæ form a rather extensive group of diurnal Lepidoptera abundantly
distributed over the whole world, and typically represented in our own country by the "cabbage,"
the "orange tip," the "brimstone" and the "wood white" butterflies. The white or yellow colour
that characterizes these familiar insects is that which recurs most frequently throughout this
group in every country; but in the tropics other forms appear by their side decorated with a
variety of colours, and assuming to some extent the forms and markings of the more brilliant
families of butterflies. It is to be observed, however, that in most cases the ground colour of the
insect is either white, yellow or black, and even in those exceptional cases in which the wings are
entirely red or bluish-grey, the female of the same species, or of one closely allied to it, presents
us with the usual simple colours. The rich rufous-brown, which so constantly reappears
throughout the great family of the Nymphalidæ, is not to be met with in a single instance in the
whole range of the present family. The metallic blue of Morpho and of the Lycænidæ, and the
rich green of various shades which occurs in most other groups of butterflies, are also entirely
absent. Although the markings are often very beautiful and very varied, well formed ocellated
spots (almost universal in the Satyridæ and very frequent in the Nymphalidæ) never once occur;
the only approach to them being the small discoidal ocelli in some species. The form of the
wings is generally rounded, rarely angular, and the hind wings are never adorned with spatulate,
linear or filiform processes, such as occur frequently in most other groups of butterflies. The
nearest approach to a tail in the whole family is found in the genus Gonepteryx, and is never
much more developed than in our own brimstone butterfly.
It appears, then, that quite independently of structural characters, the Pieridæ possess
specialities of colour, of marking, and of form of wing, which in their combination mark them
out as a natural and somewhat isolated group; and strikingly illustrate the stability of type that
sometimes obtains in what we usually deem unimportant and very variable characters. At the
same time it is to be noted that in the important structural character of the neuration of the wings,
few families exhibit so many modifications. This character serves chiefly to break up the group
into three- [[p. 302]] and-twenty genera, combined with the form of the antennæ, palpi and anal
valves, all of which vary perhaps more than is usual. The form and texture of the wings also seem
very subject to modification within certain limits, as is well seen by comparing the delicate
elongate forms of Leptalis and Leucophasia with the strong and compact Callidryas and Colias;
and in colouration no contrast can be greater than that between such genera as Euterpe and
Gonepteryx. Perhaps too there is no family more generally and uniformly distributed over the
whole earth. Pieridæ inhabit the deserts of Arabia as well as the tropical forests; they sport about
the snows of the Himalayas and roam over the icy plains of Siberia. The variety of conditions to
which they are subject is therefore as great as can well be conceived, and the considerable
generic diversity that exists among them probably indicates a great antiquity for the group, yet
the ever fluctuating characters of colour, form and marking have nevertheless their strict limits
which they in no case overpass.
The large amount of diversity in the neuration of the wings of the Pieridæ, together with the
considerable differences in the form of the antennæ and palpi, have led to the establishment of a
number of well-marked generic groups. In the "Genera of Diurnal Lepidoptera," published about
the year 1847, Mr. Doubleday admitted sixteen genera of Pieridæ. Since that time five have been
added, most of which were indicated and even named by him as distinct sections of genera. In the
present paper I add two, formed out of the old genus Pieris, thus raising the total number of
genera to twenty-three. These are distributed with tolerable equality through the great zoological
divisions of the earth; the South American and Australian regions each having eleven, the
African region ten, the varied Indian region (including the western Malay Islands) fifteen, while
the Nearctic and Palæarctic regions each possess six genera.
The extreme isolation of the South American continent from the other tropical regions is
indicated by its possessing three peculiar genera,--Euterpe, Leptalis and Hesperocharis,--while
the Indian region has but two, Prioneris and Dercas, and the Australian and Palæarctic but one
each, Elodina and Leucophasia respectively. The Nearctic and the Ethiopian regions have no
peculiar genera. Only a single genus is universally distributed over the globe, the true Pieris.
This has representatives both in the polar and the equatoreal regions, on the highest mountains
and over the hottest plains. Callidryas, Terias and Tachyris are tropical groups which range
round the equatoreal regions of [[p. 303]] the earth, but never extend far northward of the tropics.
Colias and Anthocharis on the other hand are Arctic and Alpine, occurring in both continents and
in both hemispheres, but rarely extending into tropical lowlands. In most of these widely distributed genera the species also have a wide range, and some specific forms of Callidryas,
Tachyris and Terias are almost identically reproduced in the eastern and western tropics. Most of
these are insects which roam about cultivated grounds and river banks, and collect in swarms on
wet and muddy shores; while the more restricted species and genera, such as Euterpe, Leptalis,
Leucophasia, Elodina, Thyca and Eronia, are inhabitants of woods and forests, and become
quickly exterminated by the advance of cultivation.
The following table exhibits the distribution of the genera of Pieridæ throughout the six
zoological divisions of the earth, with the number of species of each described up to this time.
[[p. 304]] By a consideration of the facts presented by the distribution of Birds and
Mammals, it has been proved that the Malay Archipelago really belongs to two primary
zoological divisions of the earth. A line drawn on the east side of the Philippines, curving to the
west of Celebes, and passing between the islands of Baly and Lombock, will divide the Indian
from the Australian region. Mr. Pascoe however is of opinion that this division will not hold
good for insects, as he finds that in Coleoptera, and especially in Longicornia, New Guinea and
the Moluccas have a much greater resemblance to Borneo and Malacca than they have to
Australia. If this statement expressed the whole of the facts of the distribution of these insects, it
would undoubtedly tend to prove that no very general causes have determined the distribution of
organic life, and that we could therefore not hope to mark out such zoological regions of the
earth as would be alike useful to the students of every branch of natural history. But I believe that
along with this general resemblance of the Coleoptera from the two extremities of the
Archipelago, there is also a diversity that points to the same division that is so strikingly
manifested in the distribution of the higher animals. As examples I may mention the great and
interesting Longicorn sub-family of Tmesisternitæ confined to the Australian region, although,
owing to unfavourable conditions, almost absent from Australia itself. Cyphogastra, which
Deyrolle considers one of the best marked genera of the family of Buprestidæ, and Eupholus
among the Curculionidæ, are similarly restricted; while the metallic Elateridæ (Campsosternus),
the giant Buprestidæ of the genus Catoxantha, and the splendid phytophagous Sagræ, are
bounded in the western half of the Archipelago by the line above indicated. Such cases as these
(and I feel sure they are much more numerous than is yet believed) would show that even the
distribution of the Coleoptera is largely influenced by the same general causes which have more
powerfully affected other groups; and I believe the cause of the anomaly presented by this order
is to be found in the fact, that, in the grand and luxuriant forests which cover the whole
Archipelago, almost all the Coleoptera are more or less intimately associated with dry or
decaying timber. The larvæ of all the most prominent groups are lignivorous, and even the
Carabidæ mostly dwell under dry or rotten bark. This circumstance is evidently in the highest
degree favourable to the diffusion and intermingling of the productions of the different islands,
and as the general physical [[p. 305]] conditions are almost identical throughout, it is not
surprising that an original diversity should have been reduced to an approximate uniformity of
type in the two regions.
If this is the true explanation of the phenomenon, we ought to find the distinction of the two
regions more marked in those orders and families where there is less probability of involuntary
diffusion. The butterflies are almost entirely free from the chance of being carried from island to
island by floating timber, but they are at all times subject to be blown great distances by storms
and by the strong and changing monsoons. This last cause of diffusion is perhaps quite as
powerful as the former one, but it is probable that these insects are not able to establish
themselves in new countries with so much certainty as the Coleoptera, (as might be expected
from the dependence of many of the species on particular plants, which they might not find in the
place to which they were carried); for we find many instances in which genera and sections of
genera are strictly limited to one-half of the Archipelago or the other. For example, the genera
Mynes and Hypocista are confined strictly to the Australian region, and Drusilla is only
represented out of that region by one species in Java. To this region also belong the genus
Elodina, the Erectheus and Priamus groups of Papilio, the Rachel group of Pieris, the Celestina
group of Tachyris, and the Mysis and Philyra group of Thyca. On the other hand, Prioneris,
Dercas and Thaumantis range over the whole Indian region, but do not cross the dividing line;
while Thestias, the Coronis group of Pieris, and the genera Euripus, Clerome, Amathusia,
Zeuxidia and Discophora, only pass over the boundary into Celebes, or along the closely linked
chain of islands east of Java to Timor. In most of these cases, the same groups that inhabit the
Moluccas and New Guinea are found also in Australia, and in some of the islands of the Pacific;
while, notwithstanding the similarity of physical conditions, and the narrowness of the dividing
seas, they do not pass into Java, Borneo, or the other western islands. These facts go to prove,
that notwithstanding the causes favourable to migration and intermixture in the case of insects,
yet the isolation of groups dependent on ancient and permanent features in the physical
geography of the country may still be very clearly traced; and if so, the primary zoological
divisions of the earth, founded on the facts of distribution presented by the higher animals, will
be found equally applicable to the objects of the Entomologist's study, since it will enable him
[[p. 306]] to account for the phenomena of distribution as he finds them to exist.
Zoological divisions of the earth's surface can only be true and useful ones, in so far as they
agree with the most ancient and permanent natural barriers to the diffusion of species. Those
divisions marked out by diversity of physical character or of climate are less important, because
they are necessarily less defined, less general in their effects, and have most likely fluctuated
much during short geological epochs. From some points of view the productions of the pampas
of Buenos Ayres may differ more from those of the Brazilian forests than do these latter from
those of the forests of tropical Africa, yet all will admit that the latter is a primary, while the
former is a secondary zoological division. However large the number of groups which can be
adduced as common to the equatoreal regions of Africa and America, but which do not extend to
the open plains of temperate South America, we should none of us admit the propriety of forming
for those groups a new division of the earth, in which the tropics of America and Africa should
form a single region. All we can do is to endeavour to account for the anomaly, and try to
discover what special causes have affected the distribution of these particular groups in a manner
very different from that of almost all the other productions of nature. The case we are
considering is one almost exactly analogous to this. The distribution of mammals, of birds, of
reptiles, and many facts in the distribution of insects, clearly indicate that the mass of land
forming Australia is but a portion of one of the primary zoological divisions of the globe,
although in it some of the peculiarities of that division reach their maximum, while others are
better exhibited in New Guinea and the Moluccas. The peculiar climate and physical condition of
Australia, however, have led to the development of many groups peculiar to it, while the same
causes have prevented others from becoming established in it, which have spread rapidly in the
damp wooded tropical islands to the north. This localization of forms is carried to a greater
extent in insects than in any of the higher animals; and so many extensive and remarkable groups
abound in Australia, and are to be seen in every collection from that country and from no other,
that the Entomologist may easily be led to ignore the equally striking facts that unite it to the
surrounding islands by a strong band of zoological affinity.
The numerical proportion of species in islands and groups of [[p. 307]] islands offers some interesting results agreeing with those which have been already obtained for the Papilionidæ. Of
any single island Java takes the lead in the number of species of Pieridæ (35), which is probably
owing to its having been more explored than Sumatra or Borneo. Next comes Celebes with 30
species, and further east Ceram with 24 species offers the greatest number, while Timor has but
18. Java and Timor may be considered the islands in the west and east which have the most
speciality, and the island of Waigiou, near New Guinea, is another which may be taken as
offering a good type of the characteristics of the Papuan fauna; yet these are all surpassed in
speciality by Celebes, which is situated between them, as shown by the following statement:--
||Species of Pieridæ.
| Java . . .
| Timor . . .
| Waigiou . . .
| Celebes . . .
We thus are led to consider the island of Celebes, with one or two small islands adjacent to
it, as forming a group equivalent to the much larger groups, which are well characterized by their
peculiar faunas, thus:--
When we consider the very limited area of Celebes, and the position in which it stands,
closely surrounded on every side by the other groups, and therefore liable to be colonized from
some of them by every ocean current and by every wind that blows, the amount of peculiarity
here exhibited is certainly very remarkable, and indicates that its present insular condition dates
from a very distant period. This is equally indicated by another class of facts of a more singular
[[p. 308]] In my paper on the eastern Papilionidæ (Trans. Linn. Soc. xxv. 1) I called attention
to the curious form of wing that characterizes most of the species of the island of Celebes. In the
Pieridæ it is almost equally prominent, as may be seen by comparing the following species, nine
in number, with the most nearly allied forms from the adjacent districts:--
Pieris eperia, as compared with P. coronis.
Thyca zebuda, as compared with T. Descombesi.
Thyca Rosenbergii, as compared with T. hyparete.
Tachyris Hombronii, as compared with T. lyncida.
Tachyris lycaste, as compared with T. lyncida.
Tachyris zarinda, as compared with T. nero.
Tachyris ithome, as compared with T. nephele.
Eronia tritæa, as compared with any other species.
Iphias glaucippe, as compared with I. glaucippe (Java var.).
In all these cases, comprising a large proportion of the Pieridæ peculiar to Celebes, the
species of that island have the anterior wings either more elongated or more acute, or the costa
much more abruptly curved near the base, than in the allied species from any of the adjacent
islands. Exactly the same peculiarity was found in almost every species of Papilio, and is to be
noticed also in many of the Nymphalidæ. I am not aware that any similar case has yet been
adduced of a peculiarity of form running through several distinct families, and characterizing a
single island only, or any equally limited district. The nearest approach to it is the apterous
condition of many Coleoptera of usually winged genera in Madeira, conjoined with the rather
fuller development of the wings in some other species, whose habits render flight essential to
them. We are all acquainted with Mr. Darwin's beautiful explanation of these facts, on the
principle of the preservation of useful variations, and we cannot doubt that the same principle has
acted in the case of the Celebes butterflies, although from our ignorance of all the physical and
biological peculiarities of that island we are not able to trace the process with so much certainty
as in the other case. The whole fauna of Celebes is isolated and fragmentary, and indicates a very
ancient origin and much extinction of species; and it may well be that the conditions which once
rendered this peculiarity an advantageous one no longer exist. We must ever remember that the
present condition of a fauna is but the resultant of an immense [[p. 309]] variety of causes and
changes, organic and inorganic, acting and reacting throughout all preceding ages. It is, therefore,
no objection to a theory that it does not explain everything, but rather the contrary. A true theory
will certainly enable us to understand many of the phenomena of life, but owing to our
necessarily imperfect knowledge of past causes and events, there must always remain
complicated knots that we cannot disentangle, and dark mysteries on which we can throw but a
straggling ray of light.
The Eastern Pieridæ present us with none of those wonderful cases of mimicry which
characterize the American genus Leptalis, and whose perfection enabled Mr. Bates to arrive at a
satisfactory explanation of the phenomenon. The comparatively small range of colour and
marking in the genera Eupla and Danais offer few facilities for that first accidental
approximation which is the necessary groundwork for the production of a more perfect
resemblance. Nevertheless there are a few interesting cases which I think must come under the
head of mimicry. The first is that of the Eastern species of Eronia, the females of which all
approximate, more or less, to the markings of the genus Danais. The particular circumstance that
makes it probable that this is a true case of mimicry is, that in several species a variety of the
female occurs with the base of the hind wings bright yellow, exactly corresponding to the colour
of other species of Danais. Thus in Malacca and Sumatra the ordinary dark female of Eronia
valeria mimics Danais similis, while the yellow variety is like Danais philomela. In the
Moluccas and New Guinea the dark females of Eronia argolis and Eronia iobæa are like Danais
sobrina and Danais meganira, while the yellow varieties resemble Danais cleona and D.
There seems, however, to be a group within the family of Pieridæ itself which is the subject
of mimicry. This is the genus Thyca, which comprises the most gay and variegated species, and is
very abundant both in species and individuals, which are almost entirely confined to the forests,
and have a very slow and weak mode of flight. They are, therefore, singularly parallel to the
South American Heliconiidæ, and probably like them have some special protection which
renders escape by flight less necessary to them than it is to the species of allied genera. That this
is the case is rendered more probable by the fact that I can point out four species of other genera
that resemble them most curiously in details of colouring: Prioneris thestylis has the under
surface [[p. 310]] marked and coloured almost exactly as in Thyca belladonna, and the female
resembles the same insect almost as much on the upper surface; both are found at Darjeeling.
Pieris sitana, a species peculiar to Ceylon, wonderfully resembles on the underside the common
Thyca eucharis which ranges all over India. Prioneris cornelia of Borneo, as figured by
Vollenhoven, is so like Thyca singhapura that I at first thought it might be a variety of that
species till I found by the description that it was structurally different. Lastly, Pieris læta of
Timor departs so much from the style of colouring of its allies and approaches so nearly to that of
Thyca belisama of Java, that I should almost look for an ally of the last species to be discovered
in Timor to serve as its pattern. We know so little of that island at present, and so many species
having the same distribution as T. belisama extend to Timor, or are represented there by an allied
form, that we might, quite independently of any connexion with P. læta, expect such a species to
be found there. On the whole, then, I think we have sufficient facts to show that phenomena of
mimicry, small in amount but of the same character with those that are so fully developed in
many other groups, do occur among the Pieridæ; and this is just what we might expect
sometimes to meet with in nature, for whenever a group of species possesses any amount of
special protection, even if it only leads some few of their enemies to avoid them, it will then
become advantageous for the species of other groups to be mistaken for these, and any accidental
variations that may tend to bring about a resemblance will generation after generation survive,
till further imitation ceases to be useful or is overbalanced by some concurrent disadvantage.
In a few cases what appears to be a dimorphic female occurs; the difference being however
one of colour only. In Tachyris paulina, the female (neombo, Boisd.) is either white with a pearly
gloss beneath, or yellow with the underside rich ochre. In an insect which I consider with some
hesitation to be the female of Tachyris celestina, a pure white and a rich yellow variety also
occur. In Tachyris cycinna both sexes are either greenish sulphur-yellow or rich chrome-orange.
In Tachyris zarinda the males are either clear reddish-orange or deep burnt crimson. In none of
these cases have intermediate forms ever been found, and though we have no direct proof in any
case that the two forms are continually produced from a single pair of individuals, there is every
reason from analogy to suppose that such is the fact. It is [[p. 311]] interesting to find, however,
that dimorphism, which is so strikingly developed among the Papilionidæ, should be represented
in a smaller degree in the closely allied group.
A very curious case of close resemblance occurs between two other insects, Pieris cynis and
Tachyris illana, which well illustrates the difference between resemblance produced by the mere
recurrence of a general type of colouring and that which is the result of the preservation of useful
variations, and which is termed "mimicry." These two butterflies are nearly of the same size, of a
white colour with a broad apical black border similarly scalloped within, and on the hind wings
both have a similarly-formed sub-macular dusky border. They differ, however, in the proportion
of the discoidal cells and in the structure of the antennæ and anal valves, so as to come into
distinct genera, and they are not found together, one being an inhabitant of the Malay Peninsula,
the other of the Philippine Islands. If the markings of these insects had been more varied and
peculiar, it would have furnished one case for the opponents of the explanation of "mimicry" by
natural selection. As it is, however, it well illustrates the exception that proves the rule. By far
the most general type of colouring in the Pieridæ, and which recurs in hundreds of species, is a
white ground with a black outer border, always most developed at the apex of the upper wings,
and very frequently less marked on the hind wings. It is not, therefore, surprising that among the
many slight modifications of this commonest and most simple type of colouration, two species
belonging to different genera should closely resemble each other externally. It is a case that
exactly corresponds to the very common type of colouration among the Carabidæ,--black elytra
with two or four rufous spots,--which recurs so frequently in many distinct genera of that
family. Such cases serve to show us how completely inadequate is the law of "reversion to a
common type" to explain the resemblance, through all the minute details of a complicated and
unusual style of marking and colouration, which is often found in cases of "mimicry."
Although the differences of neuration offer excellent characters for the establishment of
genera, it is of too uncertain and variable a nature to serve as a complete test of their affinities to
each other. I have therefore endeavoured to arrange the genera in a natural series rather by the
form of the antennæ, which seems less liable to sudden and abnormal variations, and by other
general characters; commencing as usual with the long and slender antennæ of the weak-flying
Leptalis and Euterpe, and ending with the thick [[p. 312]] antennæ of the strong-winged
Callidryas. The chief difference from the usual arrangement is in placing Terias before Pieris
and after Felder's new genus Elodina, which has hitherto been divided between Pieris and
The portion of my work to which I have devoted most time and attention has been the proper
determination of the sexes in many species of Pieris which have hitherto never been properly
located, and the separation of the heterogeneous mass of species in that genus into natural
groups. As a first step I went carefully through the whole of my collection and determined the
sex of every specimen. This was a matter of some difficulty, as there are no prominent sexual
characters in the group, and I soon found that differences of colouration and in the form of the
wings were very deceptive and had led to many errors. A little examination however soon
convinced me that the presence of anal valves in the male, almost as well marked as in the genus
Papilio, offered a certain test of sex in Pieris and its allies, and as my collection contains good
series of many of the species from different localities, I was soon led to satisfactory results in
many cases. At the same time I made the important observation that an extensive series of
species, evidently all closely allied, were characterized by large tufts of stiff hairs or bristles
springing from the base of each anal valve and often extending beyond their extremity. Both
African and American species closely allied in form and colouration had the same character,
which moreover was not found to exist in any other group of Pieridæ. The character is not one
which has been hitherto used in characterizing genera, but as its presence is in every case easily
ascertained and it groups together species which are evidently closely allied, it is probably of
physiological importance; and as it enables us to break up into natural divisions a very unwieldy
mass of species, I have determined to form the group so characterized into a genus which I term
Tachyris, as they have most of them pointed wings and a very rapid flight. Pieris panda, Godt.,
and Pieris nero, Fab., may be considered the types.
The genus Thyca, established by Wallengren for the species of Pieris which have only one
precellular subcostal nervule (and previously indicated as a section by Doubleday), is not
adopted by Vollenhoven in his Monograph of Malayan Pieridæ, because he thinks it conflicts too
much with other affinities. A careful examination of all the specimens having this character has
led me to the opposite conclusion, since the fifty-seven species I place in the [[p. 313]] genus
Thyca form, with very few exceptions, a series whose affinities for each other are very clearly
marked. There remains another small group of species which do not harmonize well with any of
the other forms, and which have been generally scattered widely through the genus. The bond of
affinity which brings them together is to be found in the costal margin of the fore wings in the
males being distinctly toothed along its entire length, forming a saw-like edge often visible to the
naked eye. The species which possess this peculiarity, now nine in number, are found to agree
closely in every other point of general structure as well as in the form and texture of the wings
and in their system of colouration; and I am informed by Mr. Watson that they have another
striking peculiarity in being quite devoid of the scales termed plumules, which are present in the
males of all other members of the old genus Pieris. Such a combination of characters appears to
me to indicate generic difference, and I accordingly propose to form for them the genus
Prioneris, of which the type will be the Pieris thestylis, Doubl.
There still remains in the genus Pieris, of which we may consider our P. brassicæ to be the
type, a considerable number of species which agree very closely in structure, though they will
range under several types of form and colouration. Even from these, however, two genera have
been separated, neither of which I here admit. The genus Aporia, Hubn., has been adopted by
some European Entomologists for Pieris cratægi and its allies, and it undoubtedly is a very
distinct form from its European congeners. But when we see that, by means of P. hippia, Men.,
and P. soracta, Moore, it is allied to such different-looking insects as P. agathon, Gray, and P.
nabellica, Boisd., which evidently form the fragments of an extensive natural group of species
not differing in any important structural characters from the true Pieris, we comprehend that it is
more by isolation and a specialised facies than by any generic peculiarity that these insects differ
from their nearest allies. The only character that can be so considered, the short palpi of P.
cratægi, is not constant in the allied forms. These constitute an Alpine and Arctic group of
Pieris, and our common species is a straggler beyond the range of its closest relations.
Wallengren has separated the African Pieris severina, Godt., and its allies, under the name of
Pinacopteryx, but the generic character he has given would apply to many other species. The
only character by which these seem to me to differ from true [[p. 314]] Pieris is in having the
anal valves of the male more or less spined. The same character, which is not mentioned by
Wallengren, is found in P. mesentina, Godt., P. coronea, Cram., and allies; the South American
P. monuste, P. ausia, and others, also have it, and it reaches its maximum in P. lycimnia, Cram.,
in which the long acute curved spines are very conspicuous; but the character is quite an isolated
one and varies too much in degree and is too difficult of determination in many cases to serve as
a chief generic character. I therefore keep these species as a section of true Pieris.
My divisions of the genus Pieris of authors may be tabulated as follows:--
A. One branch of the subcostal nervure before the end of the cell . . . THYCA (2).
AA. Two branches of the subcostal nervure before the end of the cell.
B. Males with tufts of hairs or bristles
at the anal valves . . . TACHYRIS (3).
BB. Males with the anal valves bare.
with the costa serrated . . . PRIONERIS (4).
with the costa smooth . . . PIERIS (1).
Of the external affinities of the Pieridæ it is impossible to say much. They have no direct
relation to any other family than the Papilionidæ, and even with those there is no good
connecting link. The European genera Zegris and Thais are perhaps the nearest, since they not
only offer some approximation in the structural characters of the imago, but agree also to some
extent in their metamorphoses, the larvæ of both genera enclosing themselves in a delicate web.
In the following list of species I have endeavoured to arrange them in a natural series, and
have described the new forms chiefly by comparison with well known types. I have not thought it
necessary to burden the Catalogue with many synonyms, but have in most cases referred only to
Boisduval's "Spécies Général des Lépidoptères" and to Vollenhoven's Monograph for the
species mentioned by those writers, except in cases in which there has been confusion of sexes,
varieties, or general synonymy. To avoid confusion between two works with almost identical
titles, I quote the first (quarto) Catalogue of the East India Company's Museum, published in
1829, as, "Horsfield, Cat. Lep. E. I. C.;" and the second (octavo) Catalogue of 1857, as, "Moore,
Cat. Lep. E. I. C." To make the list of described species in the [[p. 315]] family as complete as
possible I have introduced all those described by Boisduval in the "Bulletin" of the French
Entomological Society, by Lucas in the "Revue et Magasin de Zoologie," and by Felder in the
"Voyage of the Novara," although I have not been able to examine several of them, and have had
to some extent to guess at their true position and affinities.
I have drawn out a table (inserted at the end of the Catalogue) showing the distribution of
each species, as far as yet known, in the island groups and regions, and in which I have linked
together by brackets the closely allied species which may be considered as "geographical forms"
or "local varieties" by many Entomologists. Other tables show the distribution of the genera in a
For convenience of reference I add a list of the new species described, and of the old ones
whose names have been altered.
1. Pontia dione, n. s.
2. Elodina bouruensis, n. s.
3. Elodina signata, n. s.
4. Terias australis, n. s.
5. Terias ingana, n. s.
6. Terias sinta, n. s.
7. Terias rubella, n. s.
8. Terias fimbriata, n. s.
9. Terias silhetana, n. s.
10. Terias diversa, n. s.
11. Terias celebensis, n. s.
12. Terias virgo, n. s.
13. Pieris mentes, n. s.
14. Pieris narses, n. s.
15. Pieris jæl, n. s.
16. Pieris naomi, n. s.
17. Pieris tamar, n. s.
18. Pieris corva, n. s.
19. Pieris copia, n. s.
20. Pieris amba, n. s.
21. Thyca pandemia, n. s.
22. Thyca parthenope, n. s.
23. Thyca ninus, n. s.
24. Thyca singhapura, n. s.
25. Thyca ennia, n. s.
26. Thyca philotis, n. s.
27. Thyca echo, n. s.
28. Thyca hippodamia, n. s.
29. Thyca orphne, n. s.
30. Tachyris clavis, n. s.
31. Tachyris abnormis, n. s.
32. Tachyris urania, n. s.
33. Tachyris amarella, n. s.
34. Tachyris cynisca, n. s.
35. Tachyris panthea, n. s.
36. Tachyris corinna, n. s.
37. Tachyris galba, n. s.
38. Tachyris bouruensis, n. s.
39. Prioneris Vollenhovii, n. s.
40. Idmais fulvia, n. s.
41. Thestias venatrix, n. s.
42. Thestias pirenassa, n. s.
43. Thestias dissimilis, n. s.
44. Iphias borneensis, n. s.
45. Callidryas phlegeus, n. s.
NEW NAMES GIVEN TO DESCRIBED SPECIES.
Thyca pyramus = Pieris
thisbe, Gray (nec Cramer).
Tachyris alope = Pieris amasene, Boisd. (nec Cramer).
Tachyris Lucasii = Pieris pandione, Boisd. (nec Hübner).
Thestias Vollenhovii = Thestias balice, Voll. (nec Boisd.).
SPECIES OMITTED AS UNRECOGNIZABLE.
Papilio perimale, Donovan. (See Pieris
Papilio clytie, Donovan. (See Pieris teutonia.)
Papilio melania, Fab. (See Tachyris ega.
A small group of delicate eastern and tropical butterflies, generally placed next Leucophasia,
with which, however, it seems to have no near affinity. These insects fly about slowly in woods
and shady places, keeping near the ground and often settling on leaves and flowers. The species
are all very closely allied to each other. They are absent from the Moluccas and New Guinea, but
extend through the islands east of Java to Timor, and thence into North West Australia.
[[p. 317]] 1. Pontia nina, Fabricius.
Papilio nina, Fab. Ent. Syst. III. i. p. 194. Pontia nina, Bd. Sp. Gén. p. 431; Voll. Mon. Pier.
Hab.--Malacca, Sumatra, Borneo, Java, Baly, Flores, Timor (Coll. Wall.); Philippine Is. (B.
M.); India, Ceylon (B. M.).
This species varies but little throughout this extensive range. The specimens from the
Philippine Islands are very large.
2. Pontia Crookera, MacLeay.
Pontia Crookera, MacLeay, King's Survey, p. 458; Bd. Sp. Gén. p. 431.
3. Pontia niobe, Wallace.
Pontia niobe, Wall. Proc. Zool. Soc. 1866, p. 357.
Hab.--Formosa (Coll. Wall. ex Swinhoe).
Near P. nina, but a sufficiently distinct local modification.
4. Pontia dione, n. sp.
Much larger than P. nina, discoidal spot similar but larger, apical spot wanting, apical and
basal margins somewhat dusky. Beneath with the irrorations browner than in P. nina, and
extending more over the upper wings, which have an apical dusky patch. Antennæ long, tipped
with dusky orange.
Expanse 2 - 2 1/8; inches.
Hab.--Macassar (Coll. Wall.).
5. Pontia medusa, Cramer.
Papilio medusa, Cr. 150 F. Pontia medusa, Bd. Sp. Gén. 433.
A large and finely coloured species according to Cramer's figure.
6. Pontia lignea, Vollenhoven.
Pontia lignea, Voll. Mon. Pier. p. 4, pl. 2, f. 1 a, 1 b, ♂.
♀ Above dusky white, costal margin dusky irrorated; a discoidal spot as in P. nina, and a
small indistinct one above it within the apex; hind wings with a narrow dusky margin. Beneath
dusky and thickly irrorated, with a greenish tinge on the apex of [[p. 318]] the upper and over the
whole surface of the hind wings; discoidal spot on uppers dusky black. Antennæ orange-tipped.
Expanse 1 5/6 inch; wings more elongate than in P. nina.
Hab.--Menado, Celebes (Coll. Wall.).
A very distinct species, with more elongate wings than its allies.
Voy. Novara, Lep. p. 215.
This is a group of small delicate white and black butterflies, which have a peculiar type of
neuration, there being only one subcostal nervule before the end of the cell, and the lower radial
is situated so high up as to appear like a branch of the subcostal at the end of the cell. The type
has been considered a Terias, and others have been placed in the genus Pieris. They seem allied
to both Pontia and Terias, and are the only well marked group of Pieridæ which are entirely
confined to the Australian region. The species yet known are all very closely allied to each other.
1. Elodina egnatia, Godart.
Pieris egnatia, Godt. Enc. Méth. ix. p. 138. Terias egnatia, Bd. Sp. Gén. p. 678.
Hab.--N. Celebes (Wall.); Amboina (Bd.); Aru Islands (Wall.); Timor (Wall.).
The Timor specimen is smaller and has the dusky mark beneath the uppers reduced to a
subquadrate spot, but I do not like to separate it.
2. Elodina hypatia, Felder.
Elodina hypatia, Feld. Voy. Novara, p. 216.
Hab.--New Guinea (Coll. Wall.).
Distinguished by the upper wings having beneath a black apex enclosing a white stripe; but
in some specimens these markings are almost or quite obsolete, and they then only differ by the
costa above being more broadly dusky at the base.
3. Elodina therasia, Felder.
Elodina therasia, Feld. Voy. Novara, p. 215.
"Above, like E. egnatia; beneath, has the apex beyond the dark patch dusky." I have not seen
[[p. 319]] 4. Elodina bouruensis, n. sp.
Above, has the costal and outer margins broadly black with a dilated apex, and the base
broadly dusky; beneath, a curved blackish band across the apex is continued along to near the
base of the costal margin; the rest as in E. egnatia.
Hab.--Bouru (Coll. Wall.).
A sufficiently distinct species, if any of the forms of E. egnatia are to be separated.
5. Elodina angulipennis, Lucas.
Terias angulipennis, Lucas, Rev. et Mag. de Zool. 1852, p. 431. Pieris pallene, Hew. Ex.
Butt. vol. i. Pieris II. f. 8, 9.
Very distinct by the band of spots across the hind wings beneath.
6. Elodina padusa, Hewitson.
Pieris padusa, Hew. Ex. Butt. vol. i. Pieris II. f. 10, 11.
Comes near E. egnatia, from which it differs chiefly in form.
7. Elodina parthia, Hewitson.
Pieris parthia, Hew. Ex. Butt. vol. i. Pieris II. f. 12, 13.
8. Elodina signata, n. sp.
Above: upper wings black, with a somewhat irregular ovate central white patch; lower wings
white, inner margin with a black border, outer margin finely dusky and with the ends of the
Beneath: the apex of the uppers white, the ovate patch yellowish, the base slightly dusky;
lower wings white, dusky at the base, with a transverse band of dusky lunules between the cell
and the outer margin. In another specimen, without abdomen (? ♂), the lower wings immaculate
Expanse of wings 1½ inch.
Hab.--New Caledonia (B. Mus).
[[p. 320]] TERIAS, Swainson.
These small white and yellow butterflies are generally distributed throughout the tropics,
where they frequent gardens and plantations and the skirts of the forests rather than their deeper
recesses. They also assemble on the margins of streams and on the sea beach. These habits lead
to their being frequently carried off by winds, and it is thus perhaps that some of the species have
so wide a range and offer such perplexing variations. From their great similarity, and their
commonness in the tropics, they have been somewhat neglected, and we do not yet possess
materials sufficient to determine the limits of many of the forms. I have described a few new
species which appear to be sufficiently distinct, but the whole genus is at present in a very
1. Terias harina, Horsfield.
Terias harina, Horsf. Cat. Lep. E. I. C. p. 137; Bd. Sp. Gén. p. 668.
Hab.--India, Malacca, Java, Borneo, Philippines, Celebes, Batchian, Ceram, Aru Islands
This species varies little; the specimens from the Moluccas have the apical border narrower
than in those from the western islands. The female is much paler and sometimes has a broader
apical border. The Philippine form has a narrow border to the hind wings, and might almost be
considered a distinct species.
2. Terias angulata, Wallengren.
Terias angulata, Walleng. Wien. Ent. Monats. iv. p. 34.
"Alis posticis angulatis; omnibus albis, margine exteriore late nigro, introrsum undulato,
maculaque flava ad angulum anticum alarum posticarum notato, basi in parte superiore alarum
anticarum pone cellulam albo-flavescente." (Walleng.)
3. Terias læta, Boisduval.
Terias læta Bd. Sp. Gén. p. 674.
Hab.--N. W. India, Afghanistan, Nepaul, Bhotan (B. M.).
4. Terias venata, Moore.
Terias venata, Moore, Cat. Lep. E. I. C. p. 65, pl. IIa. f. 2, ♂.
Hab.--N. W. India.
The female scarcely differs, except in being more dusky.
[[p. 321]] 5. Terias santana, Felder.
Terias santana, Feld. Voy. Novara, p. 211.
Hab.--Bengal, Philippine Islands.
This appears scarcely to differ from P. venata, Moore. The female described by Dr. Felder
seems undoubtedly to be my P. vagans. I was about to describe the Philippine species as distinct,
but find that it agrees minutely with Dr. Felder's description. It may nevertheless not be the
same. The wings are rather less acute than in T. venata, and the fringe is narrower and more
6. Terias vagans, Wallace.
Terias vagans, Wall. Proc. Zool. Soc. 1866, p. 357; Feld. Voy. Novara, p. 211, ♀ (T.
Hab.--N. India, Formosa (Coll. Wall, ex Swinhoe).
7. Terias smilax, Donovan.
Papilio smilax, Don. Ins. N. Holl. pl. 20. Terias smilax, Bd. Sp. Gén. p. 660.
Hab.--Australia, Moreton Bay (Coll. Wall.).
Closely allied to T. vagans and T. venata.
8. Terias herla, MacLeay.
Pieris herla, MacLeay, King's Survey, p. 460; Bd. Sp. Gén. p. 660.
Hab.--Australia (Coll. Wall.).
A very distinct species by its reddish under surface; the specimens vary somewhat in outline,
some having the apex more rounded, and the black border terminating less abruptly, but they can
hardly be considered distinct. One specimen in the B. Mus. has almost lost the red tint of the
9. Terias australis, n. sp.
Form and size of T. smilax, but the wings more elongate, fringe yellow instead of red.
Above: upper wings as in T. smilax, but the border reaches the outer angle. Lower wings
with a black border, scalloped [[p. 322]] within, widest at the outer angle and vanishing before
reaching the anal angle.
Beneath: rich yellow; upper wings with two minute discoidal dots, and marginal dots as in T.
smilax; lower wings scantily speckled with black scales, a discoidal and a submarginal small
black spot towards the base, and very faint transverse dusky bands as in T. smilax.
Hab.--Australia, Moreton Bay (B. M.).
10. Terias ingana, n. sp.
The second subcostal branch and the disco-cellular nervure of the lower wing arising close
together as in T. herla. Wings elongate, uppers with a black border at the apex not quite reaching
the outer angle, above which are two rounded indentations; lower wings with a row of marginal
dots, sometimes connected by a narrow black edging; a black mark sometimes appears at the end
of the cell. Beneath: upper wings with a black discoidal spot and minute marginal dots; lower
wings with discoidal and basal spot as in T. australis, sometimes another within the cell, and a
transverse curved dusky band, sometimes very distinct, at others almost obsolete. Black points at
the ends of the nervures. Fringe of the upper wings dusky, of the lower yellow. Female paler.
Expanse of wings 1½ inch.
Hab.--Australia, Sidney (B. M.).
11. Terias sinta, n. sp.
Form of T. smilax; border of upper wings narrower, entire and slightly scalloped within, not
reaching the outer angle; lower wings with minute dots at the end of the nervures. Beneath:
uppers with a very faint discoidal spot; lower wings faintly black-speckled, and with spots and
band as in T. australis, more or less faintly indicated. Fringe yellow; no black marginal dots
Expanse 1½ inch.
Hab.--Australia, Moreton Bay (B. M.).
12. Terias senna, Felder.
Terias senna, Feld. Voy. Novara, p. 212.
[[p. 323]] 13. Terias drona, Horsfield.
Terias drona, Horsf. Cat. Lep. E. I. C. p. 137, t. 1, f. 13; Bd. Sp. Gén. p. 675.
Hab.--Java, Flores, Timor, Bouru, Ternate, Celebes, Ceram (Coll. Wall.).
14 Terias lerna, Felder.
Terias lerna, Feld. Sitz. Akad. d. Wiss. 1860, p. 449; Voy. Novara, p. 212.
Hab.--Amboina, Australia (Wall.); India (Feld.).
This species is exactly like T. drona above, but differs considerably on the under surface.
15. Terias rubella, n. sp.
Above, like T. drona, but the wings are shorter, and the outer margin of the uppers slightly
concave, the black border of the hind wings narrower, toothed at each nervure, not reaching the
anal angle, the fringe and two spots on the costa near the apex brick-red. Beneath, marked like T.
drona, but the whole costal margin and the fringe clear brick-red. Club of antennæ beneath, palpi
and head also red.
Expanse of wings 1¼ inch.
Hab.--Calcutta (Coll. Moore), Darjeeling, China (B. M.).
This pretty little species is quite distinct from T. drona in form, although closely resembling
it in markings.
16. Terias fimbriata, n. sp.
Form of T. hecabe. Above, clear yellow; a black border at the extremity of the uppers from
the end of the first subcostal nervule to the outer angle, pretty regularly toothed within and
dilated into a small apical patch, in which is a dusky indentation. The lower wings have only a
black spot at the end of each nervule; fringe yellow. Beneath: the whole surface is speckled with
minute black dots, a small spot at the end of each nervure, and three or four on the costal margin;
on the uppers a brown lunule at the end of the cell, a zigzag mark in the middle, and a smaller
one near the base, and a line of three or four reddish spots between the cell and the apex; lower
wings with a large brown discoidal [[p. 324]] lunule, three small ocelli near the base, and a faint
irregular submarginal band of brown spots. Head and thorax dusky.
Expanse of wings 1 7/8 inch.
Hab.--Mussooree, N. India (Coll. Moore).
Distinguished from T. blanda by its costa not black-margined, the yellow fringe, and other
17. Terias silhetana, n. sp.
Above, clear yellow; a black border at the extremity of the upper wing, commencing abruptly
between the first and second subcostal nervules, and reaching the outer angle, not regular within
and dilated into a quadrate patch at the apex; fringe black. Lower wings with a very narrow black
edge, slightly dilated at the extremity of the nervures and with a yellow fringe. Beneath, with
blackish-brown markings, disposed nearly as in T. fimbriata, but with a rusty-brown
quadrangular patch at the apex; surface sparely dotted with dusky scales, fringe of both wings
yellow. Head and thorax dusky-yellow.
Expanse of wings 1¾ inch.
Hab.--Silhet (Coll. Moore).
A well-marked species, although closely allied to T. sari and T. blanda.
18. Terias blanda, Boisduval.
Terias blanda, Bd. Sp. Gén. p. 672.
Hab.--Bouru, Amboina, Batchian, New Guinea (Wall.); Java (Voll.); China, N. India
The black border is not quite regularly toothed or sinuated within as Boisduval describes it,
yet I believe this must be his species. The Indian specimen agrees, but has the border of the hind
wings very narrow.
19. Terias phanospila, Felder.
Terias phanospila, Feld. Voy. Novara, p. 209.
20. Terias diversa, n. sp.
Male.--Colour rich yellow; markings intermediate between T. blanda and T. hecabe.
[[p. 325]] Female.--White; base somewhat dusky, markings as in the male.
Hab.--Bouru, Ternate, Ceram, Goram, Aru Islands, New Guinea, Philippine Islands (Wall.).
The males often hardly differ from T. hecabe, but the remarkable difference in the colour of
the females seems to render it necessary to separate the form. A species from the Philippine
Islands sent me as T. alitha agrees well with this species.
21. Terias hecabe, Linnæus.
Papilio hecabe, Linn. S. N. ii. p. 763. Terias hecabe, Bd. Sp. Gén. p. 669.
Hab.--India, Sumatra, Borneo, Java (Wall.); Australia (B. M.).
Var. suava, Boisd. l. c.
Hab.--Sumatra, Flores, Timor, Matabello (Wall.); India (Bd.); China (B. M.).
The varieties of this species are infinite over its extensive range, and cannot be profitably
22. Terias sinensis, Lucas.
Terias sinensis, Luc. Rev. et Mag. Zool. 1852, p. 429.
"Near T. hecabe. Wings pale yellow; uppers with a border larger and less black than hecabe,
toothed within, with the sinus less well formed. Border returning partly up the inner margin as in
T. tilaha. Border of lower wings wider than in hecabe and tilaha. Beneath paler than above, with
a marginal row of small black dots, indistinct near the fringe, otherwise without spots or
markings. In the ♀ the inner margin is not black-bordered, and the margin of the lower wings is
23. Terias nicobariensis, Felder.
Terias nicobariensis, Feld. Verh. z.-b. Gesell. Wien. 1862, p. 480.
"Near T. hecabe."
24. Terias anemone, Felder.
Terias anemone, Feld. Wien. Ent. Monats. vi. p. 24.
"Near T. hecabe."
[[p. 326]] 25. Terias sari, Horsfield.
Terias sari, Horsf. Cat. Lep. E. I. C. p. 136.
Hab.--Malacca, Sumatra, Java (Wall.); Borneo (Moore); Flores, Timor, Australia (Wall.).
A specimen from Queensland hardly differs except in having the border of the hind wings
rather narrower, and the apical patch beneath smaller; and another somewhat intermediate form
occurs at Flores and Timor.
26. Terias eumide, Felder.
Terias eumide, Feld. Voy. Novara, p. 214, ♂.
The female differs by being of a paler yellow, and sometimes much irrorated with dusky.
Hab.--N. Celebes, Sulla Islands (Wall.). Var. Batchian (Wall.).
This is intermediate between T. hecabe and T. tilaha. The Batchian variety has the border
less abruptly sinuated.
27. Terias tilaha, Horsfield.
Terias tilaha, Horsf. Cat. Lep. E. I. C. p. 136; Bd. Sp. Gén. p. 668; Voll. Mon. Pier. p. 65
Hab.--Java (Wall.); Borneo (Voll.).
28. Terias Lorquinii, Felder.
Terias Lorquinii, Feld. Voy. Novara, p. 209; Voll. Mon. Pier. p. 65 (T. tilaha, pt.).
Hab.--Celebes, Macassar and Menado (Wall.).
A beautiful species, distinct from T. tilaha of the western islands.
29. Terias alitha, Felder.
Terias alitha, Feld. Wien. Ent. Monats. vi. p. 289.
Closely allied to T. Lorquinii.
30. Terias zita, Felder.
Terias zita, Feld. Voy. Novara, p. 210, ♂.
T. zama, Feld. l. c. ♀.
Hab.--Celebes, Menado (Coll. Wall.).
[[p. 327]] This species is hardly distinct from T. Lorquinii, as my Collection shows that great
variation occurs in this group of Terias.
31. Terias tominia, Vollenhoven.
Terias tominia, Voll. Mon. Pier. p. 66, pl. 7, fig. 4, ♀. T. tondana, Feld. Voy. Novara, p. 214,
tab. xxvi. figs. 1, 2, ♂, ♀.
Hab.--Macassar and Menado (Celebes), and the Sulla Islands (Coll. Wall.).
This very handsome species varies much in the extent of the black borders in both sexes. I
possess a female in which the wings are entirely dusky black above.
This (with several other species) has been described by both Vollenhoven and Felder, in
works dated the same year, but as that of Vollenhoven was first published and sold, I have been
obliged to adopt his names.
32. Terias celebensis, n. sp. (Pl. VI. fig. 1, ♂.)
Male.--Above, black, with a suborbicular yellow patch extending from near the costa of the
upper wings to just below the cell of the lower wings, twice sinuated towards the apex of the
uppers, elsewhere regularly curved. Beneath, yellow, with spots arranged as in T. tominia.
Female.--Dusky black, with a small subovate yellow patch across the end of the cell of the
upper wings. The lower wings with the inner margin yellow, extending in an ovate patch between
the cell and the outer angle.
Expanse of wings 1 7/8 inch to 2 1/8 inches.
Hab.--Macassar, Menado, Sulla Islands (Wall.).
This beautiful and very distinct species forms an interesting transition to the male of T.
candida. It is restricted to the Celebes district, which includes the Sulla Islands.
33. Terias candida, Cramer.
Papilio candida, Cr. 331 A, ♀. Terias candida, Bd. Sp. Gén. p. 673; Voll. Mon. Pier. p. 68.
Hab.--Amboina, Ceram, Bouru, Ké Island (Coll. Wall.); Timor? (Voll.).
This species is well distinguished by the abdominal margin being black or dusky in both
sexes. Vollenhoven says it is found [[p. 328]] in Sumatra and also in Celebes; the former locality
is so improbable that I unhesitatingly reject it in the absence of direct evidence, and as to the
latter perhaps a specimen of T. celebensis has been mistaken for it.
34. Terias puella, Boisduval.
Terias puella, Bd. Voy. Astrol. p. 60, pl. 2, fig. 8, ♂; Sp. Gén. p. 674 (candida, var.).
Hab.--Batchian, Gilolo, Waigiou (Coll. Wall.).
The female resembles the male, but has the border rather wider, and is of a pale
sulphur-yellow above. In the male the black border is more regularly curved towards the costa
than in T. candida, ♂. Vollenhoven (Mon. Pier. p. 69) says his specimen of a female from
Ternate is white. This would agree with my T. virgo. Is it not an error of locality, as he does not
say that the specimen was obtained from any recent collector or other trustworthy source, and
many Ternate vessels visit the Aru Islands, the habitat of T. virgo?
35. Terias virgo, n. sp.
Male.--Above, rich yellow, the black border of the upper wings as in T. candida, of the
lower wings as in T. puella, ♂.
Female.--Colour above, pure white as in T. candida, ♀; border as in T. puella, .
Hab.--Aru Islands (Coll. Wall.).
The curious combination of characters as indicated above obliges me to separate this as a
36. Terias impura, Vollenhoven.
Terias impura, Voll. Mon. Pier. p. 70, pl. 7, fig. 5.
The form of this small species is more like that of some of the South American species of
This genus will now contain all the species of the old genus Pieris which have two subcostal
nervules given off before the end of the cell, and the males of which have neither the costa
serrated nor the anal valves tufted. The third subcostal nervule is generally well developed, but it
varies greatly, becoming very short in [[p. 329]] P. gliciria and its allies, and altogether absent in
P. daplidice and in some South American species. In P. mesentina, P. teutonia, the African P.
severina, and a few allied species, the first branch of the subcostal anastomoses with the costal,
but in others closely allied, as P. coronea, they remain distinct. As thus restricted, the genus
comprises about a hundred and twenty species distributed over every part of the globe. I divide
the Eastern species into groups characterized by peculiarities of colouration, form, or
unimportant details of structure, as follows:--
a. Coronea group.
a. Coronea group. Anal valves of male terminating in a horny point or spine; club of antennæ
rather abrupt; fore wings elongate; hind wings beneath white or yellow, with the nervures more
or less black and dilated.
b. Aspasia group. Anal valves of male ovate; club of antennæ slender; wings generally broad;
hind wings yellow or green beneath.
c. Cynis group. Wings white, pearly, delicate, broad.
d. Gliciria group. Wings rather elongate; club of antennæ rather abrupt; third subcostal
nervule very short or wanting.
e. Agathon group. Wings ample; club of antennæ ovate, flattened; palpi short; front very
hairy; third subcostal of moderate length.
1. Pieris mesentina, Cramer.
Papilio mesentina, Cr. 270 A, B. Pieris mesentina, Bd. Sp. Gén. p. 501.
Hab.--India (North and South) and Ceylon.
Some Ceylonese and Indian specimens have the female yellow-tinged above, and the lower
wings beneath of a rich ochre-yellow colour. The males do not differ.
2. Pieris teutonia, Fabricius.
Papilio teutonia, Fab. Syst. Ent. III. i. p. 199; Don. Ins. N. H. i. pl. 17, fig. 1. Pieris teutonia,
Bd. Sp. Gén. p. 473.
Hab.--Australia (Coll. Wall., B. M.).
Boisduval gives "Timor and New Guinea" also as habitats of this species; but as P. coronea
inhabits Timor, this has probably [[p. 330]] been mistaken for it. It is very possible it may extend
into New Guinea, but it seems never to have been seen there either by myself or the Dutch
Entomologists. The female is probably P. clytie of Donovan; and as it is useless to continue in
our lists species which are unrecognizable from the badness of the figures, and are probably
known under other names, I omit altogether P. perimale and P. clytie.
3. Pieris niseia, MacLeay.
Pieris niseia, MacLeay, King's Survey, App. p. 459; Bd. Sp. Gén. p. 473.
This seems to be closely allied to P. teutonia, and is perhaps not distinct. I have not seen a
4. Pieris coronea, Cramer.
Papilio coronea, Cr. 68 B, C; 361 G, H. Pieris coronea, Bd. Sp. Gén. p. 474; Voll. Mon.
Pier. p. 29.
Hab.--Java, Celebes, Baly, Lombock, Flores, Solor, Timor (Wall.).
As this species has not been found by the Dutch collectors or by myself in Sumatra or
Borneo, the above probably gives its true range. A specimen in the Brit. Mus. is labelled
"Amboina," but this is no doubt a wrong locality.
5. Pieris peristhene, Boisduval.
Pieris peristhene, Bd. Bull. Soc. Ent. 1859, p. 155.
"Above, like P. clytie ♂, but the black discoidal points larger and better marked, the marginal
row of white marks less distinct. Beneath, more like P. coronea; the hind wings black, with a row
of marginal lunules, base of the upper margin and abdominal border bright ochre-yellow. Female
smaller, of a dirty yellow-white, with a much wider black border." (Boisd.)
Expanse 2¾ inches.
Hab.--Woodlark Island (S. E. of New Guinea), New Caledonia (Coll. W. W. Saunders);
New Hebrides, Lord Howe's Island (B. M.).
In the New Caledonian insect the upper wings are white beneath, orange-tinged at the base,
with a broad outer border and a mark [[p. 331]] across the end of the cell purple-black, and a
submarginal row of white spots; the hind wings as above described.
b. Aspasia group.
6. Pieris affinis, Vollenhoven.
Pieris affinis, Voll. Mon. Pier. p. 40, pl. 5, fig. 2, ♂.
Hab.--Macassar (Wall.); N. Celebes (Voll.).
This insect bears such a curious resemblance to P. ithome ♀, that it has been always
considered to be the female of a closely-allied species. Owing to its having very small and
smooth anal valves I had always considered it to be of that sex; but having heard that Mr. Watson
had found plumules on the wing, and therefore pronounced it to be a male, I relaxed my
specimens, and by opening the valves found that he was right, and that all the specimens in my
own and other collections are males. This being the case it is evident that it has no close affinity
whatever with P. ithome, or with any other known butterfly, so that its name will be a good
example of "lucus a non lucendo," and will serve to recall the error to which its analogical
resemblance to P. ithome ♀ gave rise. It must now come in the group of true Pieris, and is nearest
to P. rachel, Bd., though forming a distinct subsection.
7. Pieris Boisduvaliana, Felder.
Pieris Boisduvaliana, Feld. Wien. Ent. Monats. vi. p. 287; Voy. Novara, pl. xxiv. fig. 8, ♂.
Hab.--Luzon, Philippines (♂, ♀, Wall. Coll.).
Evidently allied to P. rachel, but very distinct; and it is particularly interesting as showing an
approach to the curious P. affinis, which it serves to connect with this group.
8. Pieris rachel, Boisduval.
Pieris rachel, Bd. Sp. Gén. p. 469, ♂.
P. Wallaceana, Feld. Voy. Novara, p. 168, ♀.
Hab.--Waigiou (Wall.); Tidore, Ceram (Voll.); Bouru (Wall.).
These specimens agree exactly with Boisduval's description, excepting only the absence of
the small yellow spot on the under surface of the lower wings near the outer angle, of which a
trace is found only in some Bouru specimens. He gives "Java" as the [[p. 332]] locality, but that
is not to be trusted to, as collections made in other parts of the Archipelago are so frequently to
be obtained in Java. I am sorry to be obliged to cancel the name of an insect which Dr. Felder has
kindly dedicated to myself.
9. Pieris perictione, Felder.
Pieris perictione, Feld. Voy. Novara, p. 168.
This seems to be very near P. rachel. I have not seen a specimen.
10. Pieris pitys, Godart.
Pieris pitys, God. Enc. Méth. ix. p. 134; Bd. Sp. Gén. p. 470.
The wings in this species are more rounded than in P. rachel, to which it is very closely
allied. It is well described by Boisduval.
11. Pieris mentes, n. sp.
Male.--Above, like P. pitys, but with an ashy tinge on the fore wings, and generally two
spots at the apex. Beneath: the upper wings are black, with the basal half yellow-tinged and the
inner margin white, a small white mark at the end of the cell, and the apical spots (3 or 4) yellow;
the hind wings as in P. pitys.
Hab.--Lombock, Flores (Coll. Wall. and W. W. Saunders).
The dark underside of the fore wings distinguishes this very readily from pitys and rachel.
12. Pieris nabis, Lucas.
Pieris nabis, Lucas, Rev. et Mag. de Zool. 1852, p. 326, ♂. P. perithea, Feld. Voy. Novara, p.
Hab.--Australia, Moreton Bay (Coll. Wall.), Sidney and W. Australia (Coll. W. W.
A distinctly marked species, closely allied to P. rachel. A specimen in the Brit. Mus. from
New Hebrides has the fore wings more concave, and the hind wings more triangular than in the
type, and other Australian specimens vary considerably in the tint [[p. 333]] of the under surface.
This is sometimes supposed to be P. lanassa, Boisd., but it certainly does not agree with his
description. Felder's description of his P. perithea ♀ so exactly agrees with this in almost all
points that I cannot keep it separate.
13. Pieris narses, n. sp. (Pl. VI. fig. 3, ♂.)
Form of wings and markings of the upper surface exactly like P. nabis in both sexes.
Beneath: the apex of the uppers is pale rufous-brown, and the whole surface of the lower wings
earthy-brown with an orange tinge; the apical spots are obsolete, and the dusky border of the
lower wings is of a deeper colour within, forming a waved submarginal band; the base of the
upper wings is yellow-tinged, and the basal portion of the costa of the lower wings bordered with
Expanse 1 9/10 inch.
Hab.--Moreton Bay, Australia (Coll. Hewits. ♂; Coll. W. W. Saunders, ♀).
Very close to P. nabis, but appears to differ constantly in the colour and marking of the
under surface, which resembles perimale of Donovan, but the colouring of the upper surface in
his figure is so different that it cannot be considered the same species.
14. Pieris periclea, Felder.
Pieris periclea, Felder, Voy. Novara, p. 169.
Wings more elongate than P. narses, which it much resembles; the outer margin of the
Male.--Above, white; costa and base dusky, a triangular black apical patch notched in the
middle and reaching the outer angle, with a curved row of six small elongate subtriangular white
spots; hind wings with a narrow dusky border, near which the nervures are black, and a faintly
indicated dusky band within. Beneath: the apex of the uppers beyond the row of spots, which are
less distinct, rusty-ochre; hind wings entirely earthy-ochre, with a submarginal row of seven
diffused round dusky spots not far beyond the cell. The base of the uppers orange-tinged.
Expanse 1 7/8 inch.
A specimen in Mr. Hewitson's Collection agrees exactly above, but beneath the disc of the
hind wings is ochre-yellow, the sub- [[p. 334]] marginal dark spots form a continued irregular
band, beyond which is a row of indistinct whitish spots.
Expanse 2 inches.
Hab.--New Caledonia (Coll. W. W. Saunders).
These two specimens indicate a variable species, which I had at first thought distinct from
15. Pieris judith, Fabricius.
Papilio judith, Fab. Ent. Syst. III. i. p. 202. Acrea judith, Hübn. Zutr. 669, 670, ♀. Pieris
judith, Bd. Sp. Gén. p. 468; Voll. Mon. Pier. p. 21.
Hab.--Java (Wall.); (♂, ♀, B. M.).
Here commences a peculiar type of colouration, of which the present is one of the most
16. Pieris aspasia, Stoll.
Papilio aspasia, Stoll, Suppl. Cr. 33, f. 3, 3 C, ♂. Pieris aspasia, Bd. Sp. Gén. p. 469, ♂;
Voll. Mon. Pier. p. 22, ♂, ♀.
Hab.--Ceram, Amboina (Wall.).
This is the true P. aspasia of Stoll, but there are well marked permanent forms in each of the
adjacent islands, which it seems desirable to separate. Most of these offer differences in both the
sexes, and two of the females have been separately named by Vollenhoven, while he keeps the
males with P. aspasia.
17. Pieris emma, Vollenhoven.
Pieris emma, Voll. Mon. Pier. p. 24, pl. 4, f. 2, ♀.
Male.--Very like P. aspasia. Above: the fore wings have the nervures and the border rather
more broadly black, the hind wings equally rich orange, with a broader black border; beneath
there are the same differences. The female differs completely from that of P. aspasia, more so
than indicated in Vollenhoven's figure, being earthy-brown above with dull yellowish-white
spots and markings. Rather larger than P. aspasia.
Hab.--Batchian, Gilolo, ♂, ♀ (Wall.).
18. Pieris olga, Eschscholtz.
Pieris olga, Eschsch. Voy. Kotzebue, pl. 9, f. 21 a, b. Near P. aspasia. Hind wings somewhat
angular, of a lighter [[p. 335]] colour and with a broader black border; beneath, the fore wings
are much whiter, the hind wings yellow, with the costal and subcostal nervures black-bordered.
The female is intermediate between aspasia and emma, the hind wings being pale dull yellow,
with a broad irregular dusky border, and on the under side the veins and markings are
Hab.--Philippine Islands (Coll. Wall., B. M.).
A specimen from Hong Kong in the British Museum agrees with this, but is of a
reddish-orange tint on the lower wings.
19. Pieris hester, Vollenhoven.
Pieris hester, Voll. Mon. Pier. p. 24, pl. 4, f. 1, ♀.
Male.--Above: upper wings white, with a black border, but the veins much less black-edged
than in P. aspasia; lower wings clear pale orange, with a black border as in P. emma, ♂. Beneath,
differs from P. aspasia in the same degree as above.
Female.--Very distinct from P. aspasia by having the disc of the uppers yellowish-white,
with a blackish border without white or yellow spots; the lower wings dusky-orange, with an
ill-defined broad dusky border, more or less distinctly notched near the outer angle.
Hab.--New Guinea, Mysol, Waigiou (Wall.).
20. Pieris jael, n. sp.
Male.--Marked as in P. aspasia, but the nervures more clearly defined, and the hind wings
clear yellow as in P. hester, ♂, and with a black border slightly wider than in P. aspasia.
Female.--Differs in nearly the same way from the female of P. aspasia; the colour much
paler, and the markings on the under side less diffused.
21. Pieris lea, Doubleday.
Pieris lea, Doubl. Ann. Nat. Hist. xxvii. p. 23; Gen. Diur. Lep. t. 6, f. 3 (err. P. clemanthe);
Voll. Mon. Pier. p. 23, ♂ (the female as P. amalia, ♀).
Hab.--Borneo (Wall.); Banca (Voll.).
[[p. 336]] 22. Pieris amalia, Vollenhoven.
Pieris amalia, Voll. Mon. Pier. p. 23, pl. 3, f. 6, ♀ (as the male).
Male.--Above, white, the border and nervures as in P. lea, but more clearly defined and
without the obscure submarginal band; the patch on the lower wings well defined, and entirely
deep orange. Beneath, very like P. lea, but there are three yellow spots near the apex of the fore
wing, and on the hind wings the border and black nervures are narrower, and the anal region is
rich orange, shading into the pure yellow of the disc.
The female has been described and figured by Vollenhoven as the male.
Hab.--Singapore (Wall.); Sumatra (Voll.).
23. Pieris naomi, n. sp.
Male.--Above, as in P. amalia, but the orange anal patch is not clearly defined. Beneath: the
upper wings white, nearly covered by the thickened nervures and broad dusky margin, three
yellow spots at the apex and one near the costa; lower wings entirely rich orange-red, with a
rather broad purple-brown border enclosing a few reddish spots, the subcostal nervures only
Female.--Dusky brown, with whitish patches between the nervures, and a few submarginal
spots, the anal region dusky orange. Beneath: the uppers nearly as above but darker; the lower
wings purplish-brown, marked with ochre-yellow at the base and between the nervures.
Hab.--Lombock, Flores (Wall.).
A very distinct species, intermediate between P. amalia and P. temena.
24. Pieris temena, Hewitson.
Pieris temena, Hew. Ex. Butt. vol. ii. Pieris III. f. 19, ♂.
Female.--Blackish-brown, with stripes and spots between the nervures cream-white; on the
hind wings the disc is reddish-white, with the nervures dusky, and a blackish hind border with
irregular creamy spots. Beneath: the uppers are marked as in the male, but the spots are whiter;
the lower wings are less brilliant than in the male, of mixed red and yellow, the subcostal and
discoidal nervures black-bordered, and the black submarginal band more acutely toothed.
Hab.--Flores, Lombock (Wall.).
[[p. 337]] In fine specimens of the male the underside is rich orange-red. Vollenhoven (Mon.
Pier. p. 25, n.) doubts this being an inhabitant of the Archipelago, because "the Museum of
Leyden possesses a female specimen from New Zealand." I myself captured both sexes in
Lombock, and my assistant Mr. Allen did the same in Flores. I am therefore more disposed to
think that the Leyden specimen did not come from New Zealand.
25. Pieris tamar, n. sp. (Pl. VI. fig. 2, ♂.)
Form of wings as in P. temena; same size.
Male.--Above, like P. temena, but the black border about twice as wide. Beneath: differs
from P. temena by the marginal and transverse rows of spots being smaller, and the patches in
and below the cell much whiter and larger; the lower wings have a very wide black border
extending into the end of the cell, and the submarginal red spots very small, the disc coloured as
in P. temena.
Female.--Has a broad black border at the extremity of both wings, with a prolongation from
the uppers to the end of the cell. Beneath, nearly as in the male, but the spots are rather larger and
less distinctly marked.
Hab.--Baly (Coll. Wall, and W. W. Saunders).
A beautiful species, distinct in both sexes from P. temena, but more especially different in
the female. Apparently restricted to the small island of Baly, situated between Lombock and
26. Pieris læta, Hewitson.
Pieris læta, Hew. Ex. Butt. vol. iii. Pieris VII. f. 45, 46, ♂; Voll. Mon. Pier. p. 31, pl. iv. f.
Hab.--Timor (Coll. Wall. ♂).
This beautiful species evidently belongs to the present group, and seems to come best here.
The colouring of the underside is however curiously like Thyca belisama, except in the marginal
spots being red instead of orange. It may probably be a case of mimicry, like those of Prioneris
for the same group.
27. Pieris timnatha, Hewitson.
Pieris timnatha, Hew. Ex. Butt. vol. iii. Pieris VII. f. 47, 48, ♂ Voll. Mon. Pier. p. 25.
Hab.-- Tondano, N. Celebes (Coll. Wall.).
First variety.--Above: apex of the upper wings blacker, the spots scarcely marked; hind
wings without black markings on the [[p. 338]] grey border. Beneath: the costa and apex of the
uppers darker, the posterior band of the hind wings much broader, and the nervures more widely
margined, the submarginal spots very small.
Hab.--Sula Islands (Coll. Wall.).
Second variety.--Above, whiter, nearly like P. eperia, the apex dark, with small white linear
spots, and a single dark band to the end of the discoidal cell. Beneath, paler, the submarginal
band of the hind wings abruptly angular as in P. eperia, and the submarginal spots more elongate
and nearly white. Female: above, blackish, with white patches between the nervures chiefly in
and near the cell; beneath, like the male, but the nervures more broadly dark-margined.
Hab.--Macassar (Coll. Wall.).
The last variety closely resembles P. eperia beneath, but the form of the wings is quite
different. This seems to be a variable species, and a greater number of specimens would probably
present intermediate forms. The Sula Islands are zoologically a part of Celebes, as is well shown
by their birds.
28. Pieris eperia, Boisduval.
Pieris eperia, Bd. Sp. Gén. p. 470; Voll. Mon. Pier. p. 27.
Hab.--Macassar, Menado (Wall.).
This fine species differs much in form from all its allies, having the abrupt band of the costa
and the elongated wings characteristic of many Celebes butterflies. The specimens from Menado
are more richly coloured beneath.
29. Pieris pactolicus, Butler.
Pieris pactolicus, Butler, Proc. Zool. Soc. 1865, p. 455, pl. xxvi. f. 1, ♂.
Hab.--Borneo (not Bogota), (Brit. Mus. Coll.).
This is near P. coronis and P. eperia, but is quite distinct. By an oversight the habitat is
given as "Bogota," which may easily lead to its being overlooked by persons studying Eastern
30. Pieris coronis, Cramer.
Papilio coronis, Cr. 44 B, C, ♀. Pieris coronis, Bd. Sp. Gén. p. 471.
Hab.--N. India, Hong Kong (Coll. Wall., B. M.).
[[p. 339]] This species does not seem to vary much in Indian specimens, and I consider
Cramer was correct in making the allied forms distinct.
31. Pieris evagete, Cramer.
Papilio evagete, Cr. 221 F, G, ♂. Pieris coronis, part, Bd. Sp. Gén. p. 471.
Hab.--India (Coll. Wall., B. M.).
Cramer's figure well represents the male of this species; the female only differs by having
the nervures, especially the median, more dilated, and the colours rather more obscure. It is
distinguished from the female of coronis by having a row of well-marked submarginal spots on
the upper surface of the fore wings, by the yellow colour of the under surface of the hind wings,
and by their more rounded outline.
32. Pieris corva, n. sp.
Pieris coronis, var. A., Bd. Sp. Gén. p. 472, ♂.
Female.--Has the nervures round the cell of the upper wings much thickened, and the white
apical spots obsolete; beneath, as in the male.
Hab.--Java, Baly (Coll. Wall.).
This form has a well-marked character of the under surface which readily separates it from P.
coronis. It also has a posterior black border, much wider and more defined than in the allied
33. Pieris zeuxippe, Cramer.
Papilio zeuxippe, Cr. 362 E, F.
This seems to be near P. hira, Moore.
34. Pieris hira, Moore.
Pieris hira, Moore, Proc. Zool. Soc. 1865, p. 490, pl. xxxi. f. 17.
Hab.--Punjaub, Oude (Coll. Moore).
This is a small distinct species, allied to P. coronis.
[[p. 340]] 35. Pieris copia, n. sp.
Male.--Above, white; the upper wings with a black border nearly as in P. hira, but the apical
spots almost obsolete; hind wings with the spots at the end of the nervures of a greyish tint.
Beneath, the lower wings and the apex of the uppers pale greenish, the spots of the uppers as in
P. hira, but fainter; the nervures of the lower wings broadly margined with dusky olive, and a
row of small spots of the same colour parallel to the hind margin.
Expanse of wings 2¼ inches.
Hab.--Bengal (Coll. Wall. and Moore).
Closely allied to P. hira, Moore, which appears to be a small and more highly-coloured form
peculiar to West India.
36. Pieris amba, n. sp.
Male.--Above, white; upper wings with the base of the costa dusky, and a black patch across
the apex curved and crenated internally, not reaching the outer angle; lower wings with a very
faint narrow border, forming spots near the outer angle. Beneath: the uppers white, with the
costa, apex, and a partially-detached spot, pale ochre-brown; the lower wings entirely
ochre-brown, with the subcostal and median nervures and an indistinct band parallel to the hind
margin rather darker.
Expanse of wings 2 - 2 2/3 inches.
Hab.--N. India (Coll. Wall. and Moore).
Allied to P. copia and P. remba, but distinguished by the nearly uniform pale earthy-brown
colour of the underside of its hind wings.
37. Pieris nama, Moore.
Pieris nama, Moore, Proc. Zool. Soc. 1857, p. 102 (♂, ♀), pl. 44, f. 1, 2.
Hab.--N. India (Coll. Wall., B. M.).
A fine distinct species in both sexes.
38. Pieris remba, Moore.
Pieris remba, Moore, Cat. Lep. E. I. C. p. 75.
Female.--Above, blackish-brown, a large white patch on the uppers across the end of the
cell, and one on the lower wings not [[p. 341]] passing the end of the cell, and produced towards
the outer angle; base dusky. Beneath nearly as in the male. (B. M.), Ceylon.
Hab.--Canara (India), Ceylon (Coll. Wall. and Moore).
39. Pieris nadina, Lucas.
Pieris nadina, Luc. Rev. & Mag. de Zool. 1852, p. 335.
(Very near P. nama, Moore, ? same.)
♂. Upper wings about the same as P. nama; "hind wings with the extremity of the nerves
black. Beneath: uppers with a yellow band across the greenish apex; hind wings with a transverse
band of green spots.
"♀. Dusky; uppers above with two white bands, and a row of yellow spots at the tip; lower
wings also with two white bands, the hind one formed of spots placed between the nervures;
these bands more distinct beneath; nervures and margins finely black. 52 mill." (Lucas.)
Hab.--Silhet (also Java?).
Appears distinct by the description, but I have not seen a specimen.
40. Pieris nesba, Lucas.
c. Cynis group.
Pieris nesba, Luc. Rev. & Mag. de Zool. 1852, p. 324.
"♀. Fore wings, above, brown-black, with the spaces between the nervures clear grey; three
or four white marks near the apex; beneath, the same as above, with the spaces and discoidal cell
browner, a marginal row of greyish-white spots reaching the hind border. The hind wings
sulphur-yellow, with a broad black border, and marginal spots of greyish-white; beneath, lighter
yellow, with a narrower border and larger spots." (Lucas.)
A species which I have not seen, but which probably comes here.
41. Pieris cynis, Hewitson.
d. Gliciria group.
Pieris cynis, Hew. Ex. Butt. vol. iii. Pieris VIII. f. 54, ♂.
Female.--Above: upper wings blackish-brown, with a curved band of four white spots, one
large at the end of the cell, two smaller below, and one less distinct on the inner margin; lower
wings [[p. 342]] whitish, with a dusky border and base, and white fringe. Beneath, as above, but
the white marks larger and less sharply defined.
Hab.--Malacca, Mount Ophir (Wall.); Sumatra (Hewits.).
A rare species, with the aspect of an Elodina.
42. Pieris nipalensis, Gray.
Pieris nipalensis, G. R. Gray, Lep. Ins. Nep. pl. 6, f. 1, 3.
Hab.--N. India, Madras (B. M.); Himalayas, to 10,000 feet altitude (Capt. Lang).
This is the Indian form of P. brassicæ.
43. Pieris ajaka, Moore.
Pieris ajaka, Moore, Proc. Zool. Soc. 1865, p. 490, pl. xxxi. f. 16.
Hab.--N. W. Himalayas (B. M.); Lower Kunawur (Capt. Lang).
Allied to P. gliciria. Pieris melete, Ménétries, from Japan, hardly differs from this species.
44. Pieris gliciria, Cramer.
Papilio gliciria, Cr. 171 E, F. Pieris gliciria, Bd. Sp. Gén. p. 524.
Hab.--N. India, China; Himalayas, to 10,000 feet (Capt. Lang); Philippine Islands (Coll.
Wall., B. M.).
The specimens from the Philippines are rather smaller, and have the hind wings less
produced at the outer angles, but offer no definite characters to separate them specifically.
45. Pieris pigea, Boisduval.
Pieris pigea, Bd. Sp. Gén. p. 523.
I have not seen a specimen of this species.
46. Pieris daplidice, Linnæus.
Papilio daplidice, Linn. S. N. ii. p. 760. Pieris daplidice, Bd. Sp. Gén. p. 544.
Hab.--N. India, "Shipkee, alt. 13,000 feet" (Capt. Lang).
[[p. 343]] This species hardly enters the Indian region. It is truly Palæarctic.
47. Pieris kalora, Moore.
e. Agathon group. (Aporia, Hübn.)
Pieris kalora, Moore, Proc. Zool. Soc. 1865, p. 489, pl. xxxi. f. 15.
Hab.--N. W. Himalayas, 15,000 feet elevation (Capt. Lang).
Allied to P. callidice. A Palæarctic species.
48. Pieris agathon, Gray.
Pieris agathon, G. R. Gray, Lep. Ins. Nepal. pl. 8, f. 1, ♂; Bd. Sp. Gén. p. 447, ♂. P. phryxe,
Bd. Sp. Gén. p. 446; Blanchard, Voy. Jacquemont, Ins. t. 2, f. 1, ♀.
Hab.--N. India (B. M. ♂, ♀).
This curious and isolated form, I think with Mr. Moore, comes next P. cratægi and P.
soracta, which latter has the small yellow spot at the base of the hind wings beneath. The three
species agree with each other, and with the true Pieris, in all important points of structure; and I
can see no reason for separating them generically, although the short hairy palpi perhaps indicate
the affinity of this family to the Papilionidæ.
49. Pieris nabellica, Boisduval.
Pieris nabellica, Bd. Sp. Gén. p. 509.
Hab.--N. W. India, Upper Kunawur (Capt. Lang). (Coll. Moore.)
The affinities of this remarkable species were quite mistaken by Boisduval, who places it
along with P. protomedia and other African species. It is, however, clearly allied to P. agathon
and P. soracta, but is so coloured as to lead one at first sight to place it near P. eucharis, from
which group it differs in neuration and other points of structure.
50. Pieris soracta, Moore.
Pieris soracta, Moore, Cat. Lep. E. I. C. p. 84.
Hab.--N. India (Coll. Moore, B. M.).
This is evidently close to the European P. cratægi, and the [[p. 344]] appearance of the
yellow spot at the base of the lower wings beneath leads to P. agathon, which otherwise appears
(Pieris hippia, Bremer, Lep. Öst-Sibiriens, tab. iii. f. 1, from East Siberia, is very near P.
This genus is well distinguished by having only one subcostal nervure given off before the
end of the cell, the first being always absent. The body is comparatively small, the antennæ are
generally long and slender, the wings ample, the uppers often much elongated. The larvæ are
clothed with long hairs, and the pupæ are spined on the under surface.
This group is preeminently a natural one. It consists of rather large insects, which are
adorned with more rich and variegated hues than any other old-world Pieridæ; and, like so many
other richly-coloured insects, they are found only in the vast forests, which, with few
interruptions, spread continuously from the slopes of the Himalayas through Malacca and the
Malayan Islands to North Australia and the Pacific. They all fly weakly and slowly, yet they are
by no means rare, since in almost every locality I found some of the species very abundant in the
forests, flying lazily along near the ground, sometimes settling on a flower, but more generally
seeming to wander aimlessly through the pathless recesses of the forest. It is probable, then, that
they have some special protection which renders swiftness of motion and the disguise of sombre
colours unnecessary, and this seems the more probable when we find that some of them are the
subjects of mimicry, as already pointed out in the introduction, and under the genus Prioneris.
Thyca appears to be closely related to the American genus Euterpe, since it not only agrees
in many of its species having a dark ground-colour, but hardly offers any constant structural
I arrange the species under the following groups:--
a. Pasithoe group . . . Lower wings marked with a red semicircle at their base beneath.
a. Pasithoe group.
b. Belladonna group . . . Lower wings with a basal yellow patch.
c. Belisama group . . . Lower wings with a red spot at the base beneath, between the costal
and subcostal nervures.
[[p. 345]] d. Hyparete group . . . Lower wings beneath with the basal region yellow, and a
submarginal band of red, white or yellow spots.
e. Nysa group . . . Lower wings beneath black, yellow-marked at the base, and with yellow
Note.--The extensive hyparete group is further subdivided in the catalogue of species.
1. Thyca pasithoe, Linnæus.
Papilio pasithoe, Linn. S. N. ii. p. 755. Pieris pasithoe, Bd. Sp. Gén. p. 451. Papilio
porsenna, Cr. 43 D, E, ♂; 352 A, B, ♀.
Hab.--India, Silhet (B. M.).
2. Thyca henningia, Eschscholtz.
Pieris henningia, Eschsch. Voy. Kotzebue, pl. ix. f. 20 a, 20 b, ♀.
Form of T. pasithoe, but the upper wings with the outer margin concave.
Male.--Above, black: upper wings with a bluish-white band formed by a large spot in the
cell and two below it as in T. pasithoe, but larger and more defined, the outer band of spots and
that at the end of the cell absent or very faintly indicated; lower wings with a patch of rich yellow
between the cell and the abdominal margin and reaching the second median nervule; a faint
bluish-white patch across the cell to the inner margin, no submarginal spots. Beneath: the uppers
have a distinct white band across the wings, a spot at the end of the cell and four small linear
spots across the apex; the lower wings have the spots rich yellow, disposed as in T. pasithoe, but
leaving a broader black border, and the outer spots all very small; the red semicircular band is
narrower, and does not extend beyond the costal nervure to the margin of wing as in T. pasithoe.
Female.--Above, as in the male, but the band across the uppers is whiter and more defined,
and the yellow patch of the lower wings is paler and extends into the discoidal cell. Beneath it
differs in the same manner from the under surface of the male, every spot and band being the
same but rather more developed, and there are five spots instead of four in the apical band.
Expanse of wings: male 3 inches; female 3 1/3 inches.
Hab.--Manilla (Coll. W. W. Saunders).
A beautiful species, closely allied to the Indian T. pasithoe.
[[p. 346]] 3. Thyca pandemia, n. sp. (Pl. VI. fig. 4, 4 a, ♂.)
Upper wings shorter and hind wings more elongated than in T. pasithoe.
Male.--Above, black, a white spot at the end of the cell of all the wings, and at the outer
angle of the uppers, which have also the costa and a curved band of linear spots across the apex
decreasing to the outer angle within which is a linear mark, greyish-white; the lower wings have
a rich yellow abdominal patch only just passing the first median nervule, and a faint greyish band
from the inner margin across the cell. Beneath: the markings of the uppers as above but more
distinct; the lower wings have the yellow spots as in T. pasithoe, but smaller and brighter, only
the two spots between the cell and the outer row are absent, and the spot in the cell is divided
into two; the semicircular red band as in T. henningia, but still narrower.
Expanse of wings 2 5/8 inches.
Hab.--Borneo (Coll. W. W. Saunders, Brit. Mus.).
A beautiful modification of T. pasithoe, at once distinguished by the absence of the band
across the upper wings.
4. Thyca egialea, Cramer.
Papilio egialea, Cr. 189 D, E, ♀; 258 E, F, ♂. Pieris egialea, Bd. Sp. Gén. p. 450.
Hab--Java (Wall.); Sumatra, Biliton, Banca (Voll.).
Vollenhoven in his Monograph of Malayan Pieridæ (p. 7) gives also "Ceram" as a locality
for this insect, but I have no doubt this is an error, and therefore do not include it among the
localities. The whole group to which this species belongs is restricted to the Indian region.
5. Thyca crithoe, Boisduval.
Pieris crithoe, Bd. Sp. Gén. p. 450; Voll. Mon. Pier. p. 7, ♂.
Female.--Has the patch on the upper wings whiter, two spots at the end of the cell and those
at the apex large and white; on the lower wings the white patch extends only in a narrow
prolongation across the end of the cell. Beneath, differs from the male nearly as above, and on
the lower wings the yellow patch does not extend so far up the wing, leaving a notched black
band between it and the red base.
Hab.--Java (B. Mus. Coll. ♂, ♀).
[[p. 347]] The bluish patch on the disk of the upper wings is almost obsolete in some
6. Thyca thisbe, Cramer.
Papilio thisbe, Cr. 233 C. Pieris thisbe, Bd. Sp. Gén. p. 449, ♀.
Boisduval has described an Indian female as the male of Cramer's species, which I consider
distinct from that of India. I have not seen specimens.
7. Thyca pyramus, Wallace.
Pieris thisbe, G. R. Gray, Lep. Ins. Nepal. p. 8, t. 7, f. 1. (P. thisbe ♂ Bd. Sp. Gén. p. 449, is
Hab.--N. India (Coll. Wall., B. M.).
This differs so much from Cramer's figure that I believe it must form a distinct species. The
female has the lower part of the hind wings yellow and a yellow spot within the cell, and
beneath, the whole posterior border is yellow, whereas in Cramer's figure and Boisduval's
description these parts are white or dusky.
8. Thyca parthenope, n. sp. (Pl. VI. fig. 5, 5 a, ♂.)
Male.--Above: form and markings almost exactly as in T. pasithoe ♂, but the anal margin is
nearly white and the posterior row of spots less distinct. Beneath: upper wings as in T. pasithoe;
lower wings with the red semicircular band of about the same size and form as in T. pasithoe, the
markings of the rest of the wing arranged as in T. pyramus, except that the yellow colour is paler
and extends much further up the wing, filling two-thirds of the discoidal cell.
Expanse of wings 2¾ inches.
Hab.--Singapore (Coll. Wall.); Borneo (B. M.).
This insect combines the characters of two very distinct Indian species, but in the Bornean
specimen in the British Museum one of the yellow patches beyond the cell is partly divided,
showing an approach to the pasithoe form.
9. Thyca ninus, n. sp. (Pl. VII. fig. 1, ♂.)
b. Belladonna group.
Closely allied to T. pyramus, Wall. Male.--Wings broader uppers more triangular, the outer
margin not concave and pro- [[p. 348]] duced. Above: upper wings marked nearly as in T.
pyramus, but the ground colour is blacker, and the basal patches bluer and less elongate; lower
wings with the red patch at the base much smaller, not filling one third of the cell, below it a
broad transverse bluish-ashy band, paler at the abdominal margin, a large ochre-yellow patch at
the anal angle divided into four parts by the nervures; outer angle black, without whitish
markings. Beneath, almost exactly as in T. pyramus, but the red semicircle does not extend quite
so far down, and the posterior markings of the hind wing are more clearly defined and of a nearly
Expanse of wings 3 inches.
Hab.--Malacca, Mount Ophir (Coll. Wall.).
This insect differs so clearly in form, size, marking, colouration and locality from its allies,
that I have felt obliged to give it a different name, although its general appearance is such that
many Entomologists would at once pronounce it "a mere variety." Male specimens only were
taken by myself at Malacca.
10. Thyca belladonna, Fabricius.
Papilio belladonna, Fab. Ent. Syst. III. i. p. 180; Don. Nat. Repos. pl. 35. Pieris Horsfieldii,
G. R. Gray, Lep. Ins. Nepal. t. 8, f. 2.
Hab.--N. India (Coll. Wall., B. M.).
The specimens vary in size, form and colouration. Some are very black, with the spots
smaller, and scarcely a trace of yellow on the abdominal margin above.
The female is brown-black, with the spots yellowish-white. This sex seems very rare, as I
have only seen a specimen in the Collection of Mr. Hewitson.
11. Thyca sanaca, Moore.
Pieris sanaca, Moore, Proc. Zool. Soc. 1857, p. 103, pl. xliv. f. 4 (err. 6), ♂.
Hab.--Darjeeling (Coll. B. M.).
Female.--Above, pale sulphur-yellow, the dark parts as in the male, but much less diffused;
beneath pale yellow-white, the yel- [[p. 349]] low spots as in the male, the dark markings less in
extent. (B. M. Coll.)
A beautiful and rare species, allied to T. belladonna.
12. Thyca aganippe, Donovan.
c. Belisama group.
Papilio aganippe, Don. Ins. N. Holl. pl. 29. Pieris aganippe, Bd. Sp. Gén. p. 457, ♂.
Female.--Above, pale yellow; the marginal black band broader than in the male and well
defined, a row of spots between it and the cell, and a large patch at the end of the cell sometimes
connected with the border. Beneath nearly as in the male.
Hab.--Australia (Coll. Wall., B. M.).
It is difficult to locate this common Australian species. The form of its wings and the black
under surface of the hind wings, with the basal yellow and red spot, seem to indicate an affinity
to T. belladonna.
13. Thyca belisama, Cramer.
Papilio belisama, Cr. 285 A, B, C, D. Pieris belisama, Bd. Sp. Gén. p. 464; Voll. Mon. Pier.
Hab.--Java, Sumatra (Coll. Wall.).
The colour of the males varies from white to rich cream-colour, and even to orange-yellow,
and of the females from cream-colour to ochre-yellow. My Java specimens show these
14. Thyca sthenobæa, Boisduval.
Pieris sthenobæa, Bd. Sp. Gén. p. 466.
Hab.--Moluccas ("Coll. de M. Feisthamel").
Boisduval thinks this is near T. belisama. The absence of the red spot beneath is however
very remarkable, and it may perhaps be more allied to T. agostina. I have not seen a specimen,
nor is it in the Leyden Museum.
15. Thyca glauce, Butler.
Pieris glauce, Butler, Proc. Zool. Soc. 1865, p. 431, pl. xxv. f. 2, ♂.
Hab.--Borneo (B. M.).
A well-marked local form of T. belisama.
[[p. 350]] 16. Thyca Descombesi, Boisduval.
Pieris Descombesi, Bd. Sp. Gén. p. 465.
Hab.--N. India, Cochin-China (Coll. Wall., B. M.).
17. Thyca zebuda, Hewitson.
Pieris zebuda, Hew. Ex. Butt. vol. iii. Pieris VII. f. 49, 50, ♀. P. Descombesii, Voll. Mon.
Pier. p. 18.
Hab.--Celebes (Coll. Wall.).
This species is very distinct from T. Descombesi, both in form and colouration of both sexes.
The male has the excessively elongated wings so characteristic of the butterflies of Celebes.
18. Thyca aruna, Boisduval.
Pieris aruna, Bd. Sp. Gén. p. 466, ♂; Hew. Ex. Butt. vol. ii. Pieris III. f. 20, 21, 22; Voll.
Mon. Pier. p. 18. P. bajura, Bd. Sp. Gén. p. 467, ♀.
Hab.--Waigiou, Batchian (Wall.); Obi (Voll.); New Guinea (Voy. Coq.).
This is perhaps the most beautiful and remarkable of all the Pieridæ, its two sexes presenting
four totally distinct surfaces, all finely coloured.
19. Thyca harpalyce, Donovan.
Papilio harpalyce, Don. Ins. N. Holl. pl. 18. Pieris harpalyce, Godt. Enc. Méth. ix. p. 149;
Bd. Sp. Gén. p. 458.
Hab.--Australia, Moreton Bay (Coll. W. W. Saunders, B. M.).
One of the handsomest of the Pieridæ on the underside.
20. Thyca nigrina, Fabricius.
d. Hyparete group.
Papilio nigrina, Fab. Ent. Syst. III. i. p. 200. Pieris nigrina, Sw. Zool. Ill. 2 ser. pl. 69; Bd.
Sp. Gén. p. 459.
Hab.--Australia (Coll. Wall., B. M.).
A common species in most parts of Australia.
This extensive group may be subdivided according to details of colouration.
[[p. 351]] *A band of red spots, the nervures black and dilated beneath.
21. Thyca hierte, Hübner.
Papilio hierte, Hüb. Zutr. f. 77, 78. Papilio antonoe, Stoll. 33, f. 2, 2 b (nec Cramer). Pieris
antonoe, Bd. Sp. Gén, p. 454.
Hab.--China (Coll. Wall.).
Male.--Wings broader than in the type, nervures and apex more dusky. Beneath: the upper
wings have the nervures much more broadly dusky, leaving the apical spots smaller and not
yellow-tinged; the lower wings have the whole basal and abdominal region rich orange instead of
yellow, the marginal spots larger and deeper red, the two inner ones shading into the orange tint.
Hab.--Siam (Coll. W. W. Saunders); India (Coll. Moore).
Female.--Above: upper wings dusky black, with greyish lines in the cell and between the
nervures, and a band of six linear spots across the apex; lower wings with the nervures broadly
dusky, the discoidal cell and the abdominal margin nearly white. Beneath: the uppers black, with
the spaces between the nervures whiter than above, and the upper spots of the apical band
orange-tinged; the hind wings nearly as in the male, with the orange colour extending farther
towards the outer angle, and the red marginal spots all well defined.
Hab.--Burmah (Coll. Moore).
In the British Museum Collection are a series of specimens from various parts of tropical
Asia, no two of which are exactly alike, and which show that, though the extreme forms are well
marked, they are connected by such a series of completely intermediate forms that it is
impossible to define them.
22. Thyca eucharis, Drury.
Papilio eucharis, Drury, Ins. ii. t. 10, f. 5, 6; Cr. 201 B, C, 202 C (nec Fab.). Papilio
hyparete, Fab. Ent. Syst. III. i. p. 178 (nec Linn.). Pieris epicharis, Godt. Encl. Méth. ix. p. 153;
Bd. Sp. Gén. p. 456.
Hab.--North and N. W. India (Coll. Wall., B. M.).
A very common species in collections.
[[p. 352]] 23. Thyca hyparete, Linnæus.
Papilio hyparete, Linn. S. N. ii. p. 763; Clerck, Icon. t. 38, f. 2, 3 (nec Fab.). Pieris hyparete,
Bd. Sp. Gén. p. 455; Voll. Mon. Pier. p. 9. Papilio antonoe, Cr. 187 C, D, 320 A, B.
Hab.--Java, Borneo, Singapore (Wall.); Sumatra (Voll.).
This species presents slight but apparently permanent differences in each locality. Taking
those of Java as the type, the Singapore form has a narrower black border to the hind wings, the
outer red spot is wanting beneath, the three inner red spots are much larger, and the anal margin
is of a deeper yellow colour. The Borneo form has no black hind border but a broad ashy one,
while beneath it is intermediate between the preceding and the type. Both however are larger,
and have more elongate wings than the Java specimens.
Vollenhoven gives "Ceram" as another locality, but this is evidently an error.
24. Thyca luzoniensis, Felder.
Pieris luzoniensis, Feld. Wien. Ent. Monats. vi. p. 285.
Hab.--Luzon, Philippines (Coll. Wall., B. M.).
"A geographical form of P. hyparete" (Feld.). This has broader wings than the other forms,
but is hardly more distinct than those of Sumatra or Borneo.
25. Thyca Rosenbergii, Vollenhoven.
Pieris Rosenbergii, Voll. Mon. Pier. p. 11, pl. 2, f. 6; pl. 3, f. 1. P. Lorquinii, Feld. Voy.
Novara, p. 159, t. xxiv. f. 9, 10.
Hab.--Celebes, Macassar (Wall.); Menado (Voll.).
A most beautiful and distinct species, excelling all others of this sub-group as those of
Celebes so often excel their allies. Dr. Felder's name for this species is, I believe, of a later date
than Vollenhoven's, though the two publications are of the same year.
26. Thyca hæmorrha, Vollenhoven.
* * Submarginal band beneath the lower wings white; nervures black and dilated.
Pieris hæmorrha, Voll. Mon. Pier. p. 10, pl. 2, f. 5, ♂.
Hab.--Banca (Leyden Mus.).
This very interesting species shows that the little island of Banca, though separated by a
narrow strait only from Sumatra, [[p. 353]] yet possesses its peculiar species of insects as it does
of birds and mammals.
27. Thyca agostina, Hewitson.
Pieris agostina, Hew. Ex. Butt. vol. i. Pieris I. f. 1, 2, ♂.
Female.--Above: upper wings black, with ashy patches in the cell and between the nervures,
and a row of elongate white spots near the outer margin; lower wings pale ochre-yellow,
black-bordered behind, with a marginal row of ashy spots, and the nervures slightly margined
with dusky atoms. Beneath as above, but the markings on the upper wings whiter and more
defined, the lower wings bright ochre-yellow.
Hab.--North India (Coll. Hewitson and Moore).
28. Thyca blanca, Felder.
Pieris blanca, Feld. Wien. Ent. Monats. vi. p. 284.
This species seems to me nearer to T. agostina than to T. themis (with which Felder allies it),
on account of the black-bordered veins of the upper wings beneath, which I think is a good
character for arranging the species in a natural series.
29. Thyca singhapura, n. sp. (Pl. VII. fig. 2, ♂.)
Male.--Wings elongate; above, white, the costal margin and the outer half of the uppers
dusky, nearly black at the apex, and with an ill-defined inner edge; lower wings with a narrow
interrupted black border behind, within which dusky patches of scales extend a short distance
along the nervures. Beneath: upper wings white, the nervures of the upper half black-margined,
the apex blackish, leaving a row of six distinct ovate white spots; lower wings bright yellow, the
nervures black-bordered, and a rather broad black border round the hind margin enclosing a row
of six whitish spots, the inner ones bifid, the outermost yellow-tinged.
Expanse of wings 3 1/8 inches.
Hab.--Singapore (Coll. Wall.).
[[p. 354]] * * * Beneath with nervures not black or dilated, submarginal posterior band red and
white, or white.
30. Thyca argenthona, Fabricius.
Papilio argenthona, Fabr. Ent. Syst. III. i. p. 200, ♀. Pieris protochiris, Bd. Sp. Gén. p. 457,
Male.--Above, like the female, but white instead of yellow, the apical and posterior bands
narrower, margined within with an ashy tint. Beneath the same difference of tint and markings.
Hab.--Australia (Coll. Wall., B. M.).
31. Thyca peribæa, Godart.
Pieris peribæa, Godt. Enc. Méth. ix. p. 154; Bd. Sp. Gén. p. 453; Voll. Mon. Pier. p. 8.
Hab.--Java (Leyden Mus.).
This species has its affinity with T. argenthona, rather than near Prioneris philonome, where
Boisduval places it, or near T. hyparete, according to Vollenhoven. The absence of black on the
veins beneath and the white-bordered red spots indicate this.
32. Thyca themis, Hewitson.
Pieris themis, Hew. Ex. Butt. vol. ii. Pieris V. f. 31, 32, ♂.
Female.--Has the base dusky, the apical half of the uppers and a broad margin on the hind
wings black, enclosing a few faint yellowish spots; the disc of the hind wings is pale yellow.
Beneath, nearly as in the male, but with the black apex and hind margin rather wider, and the disc
of the hind wings deeper yellow.
Hab.--Philippine Islands (Coll. Hewitson).
On the whole, I consider this species nearest to argenthona, with which it agrees in having
the veins not black-bordered beneath, and a somewhat similar distribution of black and yellow on
a white ground, the posterior band of spots being white instead of red.
* * * * White with a black border in both sexes, nervures not black and dilated beneath, posterior
band yellow or white.
33. Thyca gabia, Boisduval.
Pieris gabia, Bd. Sp. Gén. p. 478, ♂; Voll. Mon. Pier. p. 38, ♀.
Hab.--Waigiou (Voy. Coquille); New Guinea (Voll.).
[[p. 355]] A rare species, which I did not myself meet with, and of which I have not seen a
34. Thyca dice, Vollenhoven.
Pieris dice, Voll. Mon. Pier. p. 39, pl. 4, f. 7.
Hab.--New Guinea, Waigiou (Voll.).
Another closely allied species of this group, not in the English collections.
35. Thyca ennia, n. sp. (Pl. VII. fig. 4, ♂, ♀.)
* * * * * Females very dark, posterior band beneath red, nervures not black or dilated.
Male.--Above, white, upper wings with the costa dusky, and a black border at the apex
regularly curved within, enclosing two small white spots and terminating in a point at the outer
angle; lower wings with a narrow hind border, not reaching the outer angle. Beneath: a row of
five apical spots, the two uppermost of which are largest and yellow, the base of the lower wings
yellow-tinged, and the hind border enclosing a line of narrow yellow spots.
Female.--Above, white, with a much broader dusky border to the anal angle, the base of the
uppers dusky, and a band at the end of the cell connecting the costa with the outer border.
Beneath, there is a band of six spots near the outer margin of the uppers, the second and third
being larger and yellow; the base of the wings, the anal margin, and several marks between the
nervures, are also yellow; the posterior band is much broader, its inner margin straight and just
beyond the end of the cell, and has a submarginal row of six horizontal yellow spots.
Expanse of wings 2 3/8 - 2 5/8 inches.
Hab.--Waigiou (Coll. Wall.).
This is very near T. dice, which seems intermediate between this and T. gabia. It is curious
that all should be found in the same small island. The upper side of T. dice is, according to
Vollenhoven's description, like my male, and the under side like my female.
36. Thyca bagoe, Boisduval. (Pl. VII. fig. 3, 3 a, ♂, ♀.)
Pieris bagoe, Bd. Sp. Gén. p. 461, ♀.
Male.--Above, white; uppers with the costal margin and apex and a band from the costa to
the middle of the outer margin, [[p. 356]] black, leaving four white apical spots formed by the
transverse nervures; lower wings white, with a very narrow border behind. Beneath: upper wings
nearly as above; lower wings rich yellow at the basal and anal margins, and a rather broad black
band behind, enclosing a row of six vermilion-red spots hardly separated by the nervures; the
second and third spots from the anal angle are much the largest.
Expanse of wings 3 inches.
Hab.--Aru Islands (Wall. ♂, ♀); New Ireland (Voy. Coquille).
A fine species, commencing a series with dark females.
37. Thyca mysis, Fabricius.
Papilio mysis, Fab. Ent. Syst. III. i. p. 200. Pieris mysis, Godt. Encyc. Méth. ix. p. 150; Bd.
Sp. Gén. p. 460, ♂.
Female.--Above: base of the wings tinged with yellow, marginal bands broader, and a row
of five spots at the apex. Beneath: basal half of the wings yellow-washed, and the two upper
apical spots yellow.
Hab.--Australia (Coll. Wall.).
38. Thyca cruentata, Butler.
Pieris cruentata, Butl. Proc. Zool. Soc. 1865, p. 455, pl. xxvi. f. 2. P. mysis, var. lara, Voll.
Mon. Pier. p. 12.
Hab.--Mysol (B. M.); New Guinea (Leyden Mus.).
39. Thyca pcilea, Vollenhoven.
Pieris pcilea, Voll. Mon. Pier. p. 13, pl. 3, f. 3, ♂.
Hab.--Gilolo, Morty (Voll.); Batchian (Coll. Wall.).
My specimens from Batchian seem so near Vollenhoven's that I do not like to separate them.
The red band beneath is broader in my insects, contains an additional small spot at each end, and
melts into the orange tint of the anal angle. The female agrees more closely.
40. Thyca candida, Vollenhoven.
Pieris candida, Voll. Mon. Pier. p. 11, pl. 3, f. 2, ♂.
Hab.--Batchian (Leyden Mus.).
Vollenhoven places this species in the hyparete subdivision, but I think its affinities are with
the mysis subdivision, on account of the [[p. 357]] absence of thickened nervures beneath the
anterior wings. It is not impossible that T. herodias from Gilolo may be the female of this, as
many of the species of these two islands are identical.
41. Thyca herodias, Vollenhoven.
Pieris herodias, Voll. Mon. Pier. p. 14, pl. 3, f. 4, ♀.
Hab.--Gilolo (Leyden Mus.).
This curious form seems quite distinct from T. pcilea, although very closely allied to it.
42. Thyca timorensis, Boisduval.
Pieris timorensis, Bd. Sp. Gén. p. 459, ♂; Voll. Mon. Pier. p. 12. P. vishnu, Moore, Cat. Lep.
E. I. C. pl. II a, fig. 5, ♀.
Hab.--Timor (Coll. Wall.).
This fine species appears to be best placed near T. mysis, from which it chiefly differs in the
great extent of the black border. It is, however, equally closely allied to T. isse.
43. Thyca philyra, Godart.
Pieris philyra, Godt. Enc. Méth. ix. p. 150; Bd. Sp. Gén. 462; Voll. Mon. Pier. p. 15. Papilio
hyparete, Cr. 210 A, B, 339 E, F (nec Linn.).
Hab.--Amboina, Ceram (Coll. Wall., B. M.).
Vollenhoven says the females from Sumatra "have no black palmated patch on the lower
wings beneath, but only the black nervures dilated." This agrees with Cramer's figure and with
my specimen from Amboina. I have no hesitation, therefore, in considering the former locality as
an error, more especially as the collector's name is not given.
44. Thyca philotis, n. sp. (Pl. VIII. fig. 4, ♂.)
Very near T. philyra, but with many points of difference, as follows:--
Male.--Above, the apical patch is much larger and the posterior border rather wider and not
so clearly defined. Beneath: the boundary of the black colour on the upper wings is in a totally
different line, passing across the end of the cell to a little above the outer angle, and leaving a
large patch of white between it and [[p. 358]] the cell; the base of the wing is purer yellow; on
the lower wings the boundary between the black and yellow portions is nearer the base of the
wing, the marginal red spots are smaller, and there is only a small detached portion in the cell.
Female.--Above: the basal portion of the wings is nearly pure creamy-white, and there is a
row of small indistinct white spots along the hind margin. Beneath: the yellow colour of the base
of the upper wings extends downwards, changing to white at the lower margin and touching the
outer angle; the lower wings are almost exactly like those of T. philyra ♂, except that the red
spots are rather broader and extend higher up near the abdominal margin. Size the same as of
large specimens of T. philyra.
Hab.--Bouru (Coll. Wall.).
This is a beautiful local modification of T. philyra in an adjacent island.
45. Thyca duris, Hewitson.
Pieris duris, Hew. Ex. Butt. ii. Pieris V. f. 34, ♂.
This rare and very distinct species comes nearest to T. philyra. The female is unfortunately
46. Thyca isse, Cramer.
Papilio isse, Cr. 55 E, F, 339 C, D. Pieris isse, Bd. Sp. Gén. p. 462; Voll. Mon. Pier. p. 14.
Hab.--Amboina, Ceram (Coll. Wall., B. M.).
An abundant species in the forests of these two islands.
47. Thyca echo, n. sp. (Pl. VIII. fig. 3, ♀.)
* * * * * * Beneath: upper wings black, lower wings yellow, nervures not dilated.
Very near T. isse, but with many points of difference, as follows:--
Male.--Above, the apical border of the upper wings is much wider, but does not extend so
far towards the outer angle, it contains two minute white dots; the posterior margin of the hind
wings is also broader and entirely black. Beneath: the disc of the upper wings is white, faintly
yellow-tinged at the base, the spots at the apex are smaller, and the black border is quite
differently shaped, curving from the costa at the end of the cell to about the middle of the outer
margin; the lower wings are two- [[p. 359]] thirds black, and one-third yellow, the division not
being clearly defined as in T. isse.
Female.--Above: basal half of the wings pale yellow, the rest black without apical spots;
beneath, the apical spots are very small, the black border of the uppers is shaped nearly as in the
male, the part near the outer angle being white; the lower wings have the black extending a little
further up, the band of orange spots very small, and the anal angle rich orange. Size the same as
Hab.--Bouru (Coll. Wall.).
This species differs from T. isse in almost exactly the same points that T. philotis differs
from T. philyra,--a very curious fact in connexion with the laws of variation, since it shows a
definite modifying influence in particular localities, which effects the same changes in distinct
species. I may remark also that in both cases the hind wings of the Bouru forms are more
elongate than those of Ceram and Amboina.
48. Thyca chrysomelæna, Vollenhoven.
Pieris chrysomelæna, Voll. Tijd. voor Ent. 1866, p. 57, pl. i. f. 12.
Hab.--Kaioa Island (Moluccas). (Leyden Mus.)
This very beautiful species will come at the head of the dorimene subdivision, and is most
nearly allied to T. echidna. It was obtained by the late Herr Bernstein in a small island, where I
had myself captured several new species of Coleoptera, but, perhaps owing to the time of year,
very few Lepidoptera.
49. Thyca echidna, Hewitson.
Pieris echidna, Hew. Ex. Butt. vol. ii. Pieris V. f. 35, 36.
Hab.--Ceram (Coll. Wall.).
I obtained only a single specimen of this distinct and beautiful species in the island of Ceram.
50. Thyca hippodamia, n. sp. (Pl. VIII. fig. 1, ♂.)
Male.--Above, white, the colours of the under surface showing [[p. 360]] through; a narrow
black border on the costal, outer, and hind margins; near the apex of the upper wings are four or
five white spots, formed by a dusky transverse band from the outer margin, which does not reach
the costa, and is crossed by black nervures; base of the wings dusky. Beneath: upper wings
uniform black, with an apical band of five white spots; lower wings rich chrome-yellow, paler on
the upper margin, with a rather broad black border, in which is a marginal row of white spots
tinged with yellow.
Expanse of wings 2½ inches.
Hab.--Aru Islands (Coll. Wall.).
A species which connects the preceding with T. dorimene, which has hitherto appeared
51. Thyca dorimene, Cramer.
Papilio dorimene, Cr. 387 C, D. Pieris dorimene, Bd. Sp. Gén. p. 464; Voll. Mon. Pier. p.
Hab.--Ceram, Amboina (Coll. Wall., B. M.).
This pretty species is not uncommon in the forests of these islands.
52. Thyca dorylæa, Felder.
e. Nysa group.
Pieris dorylæa, Feld. Voy. Novara, p. 182, ♀.
Hab.--Aru Islands (Coll. Felder).
A distinct form, allied to T. dorimene. Dr. Felder obtained it in an old Dutch collection. The
locality may therefore be erroneous, though it is a probable one.
53. Thyca momea, Boisduval.
Pieris momea, Bd. Sp. Gén. p. 477; Voll. Mon. Pier. p. 31.
Hab.--Java (Coll. Wall., B. M.).
54. Thyca nysa, Fabricius.
Papilio nysa, Fab. Ent. Syst. III. i. p. 195. Pieris nysa, Bd. Sp. Gén. p. 476, ♂, ♀.
Hab.--Australia (Coll. Wall., B. M.).
[[p. 361]] 55. Thyca lanassa, Boisduval.
Pieris lanassa, Bd. Sp. Gén. p. 477.
Hab.--Australia ("Paris Museum").
This appears to be near T. nysa. I have not seen a specimen, and it seems a rather doubtful
56. Thyca orphne, n. sp. (Pl. VIII. fig. 2, ♂.)
Form of the Aspasia group of Pieris. Male.--Above, white, with the base dusky; upper wings
with a narrow black border from the apex to near the outer angle, the costa dusky, as well as the
extremity of the cell, and a broad marginal band beyond as if smeared with black; the lower
wings have a very narrow dusky edging behind. Beneath: the uppers have the base dusky, a broad
patch at the end of the cell nearly black, and a broad dusky outer margin, enclosing a row of six
white spots, the two lower the largest; lower wings black, but nearly covered with
chrome-yellow, which covers the whole abdominal margin to the middle of the wing, beyond
which are three large oval spots near the outer angle; another band of yellow crosses the lower
half of the cell, and reaches up to the inner margin near the base of the wing.
Expanse of wings 2 3/8 inches.
Hab.--Malacca (Mount Ophir) (Coll. Wall.).
This curious species is rather difficult to locate. Its neuration shows it to belong to the genus
Thyca, and it seems best placed near nysa and momea on account of its black under wings
beneath, although the amount of yellow with which it is adorned gives it a different aspect.
57. Thyca georgina, Felder.
TACHYRIS, n. g.
Pieris georgina, Feld. Wien. Ent. Monats. v. p. 298; Voy. Novara, tab. xxiv. f. 4, 5.
Hab.--Luzon, Philippines (Coll. Felder).
Judging by the form of the wings and the style of colouration, this species is allied to T.
orphne, and comes in the nysa group.
Head moderate, palpi long, acutely pointed; antennæ of moderate length, terminating in a
slender club; thorax stout; abdo- [[p. 362]] men slender. Anal valves of the male elongated and
provided with a tuft of long and stiff hairs at each side at the base beneath, as shown in the
accompanying figures, compared with the same part in allied genera:--
1. Tachyris nathalia, Felder.
2. Tachyris nero, Fabricius.
3. Pieris agathon, Gray.
4. Pieris teutonia, Fabricius.
Upper wings with the apex acute or much produced, two subcostal nervules given off before the
end of the cell, the third of moderate length or very short, and in some species quite wanting;
upper radial as a branch of the subcostal at some distance beyond the cell. Sexes often differing
widely. Larva hairy, and with four or six longitudinal rows of spines; pupa with two lateral
This genus does not differ materially in neuration from Pieris, but the remarkable character
of the strong tufts of hair at the anal valves of the males enables us to bring together a number of
allied forms, which, whenever the male sex is obtained, can be referred to the genus with the
greatest certainty. It is restricted to the tropics, but comprises species from all the great divisions
of the globe, and is particularly abundant in the Indian and Australian regions, comprising more
than fifty species. The African species sylvia, Fab., and eudoxia, Drury, belong to this genus, and
a few others allied to them; while in South America, margarita, Hübner, and several
closely-allied species, appear to be its only representatives. In the Malay Archipelago, besides a
host of species which present the usual white or yellow tints of the family, it contains others
whose rich hues of cinnabar-red, orange, and greyish-blue, are altogether peculiar. Most of the
species fly swiftly, and many of the males assemble in troops about wet places and on river
margins, after [[p. 363]] the manner of the genus Callidryas. The Eastern species may be
conveniently arranged in the following groups:--
a. Panda group . . . Male white or yellowish, fore wings elongated acute; female
dark-bordered, wings more rounded.
a. Panda group.
b. Liberia group . . . Male bluish or dark brown, fore wings acute; female dusky, or white and
c. Nero group . . . Male red or orange, fore wings acute; female dusky.
d. Pandione group . . . Broader, colours white and black; lower wings beneath dusky
irrorated, with a spot on the disco-cellular nervule.
e. Polisma group . . . Wings broad, delicate, white and black, not irrorated beneath; discoidal
1. Tachyris Hombronii, Lucas.
Pieris Hombronii, Luc. Rev. et Mag. de Zool. 1852, p. 325; Voll. Mon. Pier. p. 5, pl. 2, f. 3,
Hab.--Northern Celebes (Coll. Wall.).
This very fine species must come into the present genus, although at first sight it resembles
Thyca hyparete and its allies. The species to which it is most closely allied is Tachyris lynceola
(which inhabits Timor), of which it may be considered a large and modified form. It possesses
the characteristic abrupt curvature near the base of the fore wings which is found in so many of
the Celebes butterflies. (See Trans. Linn. Soc. xxv. pp. 14-19.)
2. Tachyris cardena, Hewitson.
Pieris cardena, Hew. Ex. Butt. vol. ii. Pieris III. f. 17, 18; Voll. Mon. Pier. p. 25. Pieris
hagar, Voll. Mon. Pier. p. 38, pl. 4, f. 6.
Hab.--Malacca, Borneo, Sumatra (Coll. Wall.).
My specimens are intermediate between Hewitson's and Vollenhoven's figures, and I have
no doubt but that they represent [[p. 364]] one rather variable species. It is somewhat aberrant in
this group, but serves to connect T. hippo and ada with T. nerissa.
3. Tachyris nerissa, Fabricius.
Papilio nerissa, Fab. Ent. Syst. III. i. p. 192. Pieris nerissa, Bd. Sp. Gén. p. 535, ♂. Papilio
zelmira, Cr. 320 C, D, E, F. Pieris zelmira, Bd. Sp. Gén. p. 533, ♀.
Hab.--Indian Peninsula, Philippine Islands (Coll. Wall., B. M.),
The male is pretty constant and is well described by Boisduval. The female is very variable,
some specimens being nearly white, with dusky costa, apex and connecting band, and the
markings beneath, nearly obsolete; others are dusky brown, with patches on the upper and lower
wings and submarginal spots white or buffy-white, while beneath they are richly banded and
margined with chrome-yellow on a white and dusky ground. Innumerable varieties connect these
extremes. The Philippine specimens are smaller, and the posterior band of the lower wings
beneath crosses at the end of the cell. The markings of the under side of the male are sometimes
(Pieris larissa, Feld. Voy. Novara, p. 166.
Is this not amasene, Cr., from China?)
4. Tachyris lynceola, Felder.
Pieris lynceola, Feld. Voy. Novara, p. 164, ♂.
Female.--Above, blackish-brown; upper wings with an oblong white patch notched
exteriorly, situate below the cell and the second median nervule, and two small ovate white spots
(the upper much the largest) just beyond the end of the cell towards the apex of the wings; lower
wings with a large ill-defined white patch, occupying the disk of the wing and reaching the
anterior and anal margins. Beneath, pearly white, with a broad rusty-brown border, and the two
subapical spots rather larger and more diffused; there is also a faint yellowish longitudinal stripe
in the discoidal cell of the uppers.
Expanse of wings 2¼ - 2 3/8 inches.
Hab.--Timor (Coll. Wall.).
This differs very little from T. lyncida in the male, while the female is widely different.
[[p. 365]] 5. Tachyris lyncida, Cramer.
Papilio lyncida, Cr. 131 B, ♂. Pieris enyo, Bd. Sp. Gén. p. 481, ♂. Pieris hippo, var., Voll.
Mon. Pier. p. 42, ♀.
Hab.--Flores, Lombock, Baly, Java (Coll. Wall., B. M.).
The Java males vary somewhat, but are often undistinguishable from those of Baly and
Lombock. For some time I considered T. lynceola, Feld., to be Cramer's lyncida, and described
this as a new species:--
Male.--Above, like T. lynceola ♂, but the black border narrower, not enclosing the oval
white apical spot, which is larger and bounded above by the powdery white costal band; on the
hind wings the border is very narrow and vanishes before reaching the anal angle. Beneath,
differs from T. lynceola in the same way as above, and has the margin of a paler brown. Larger
and has rather more pointed wings than T. lynceola.
Female.--Varies much, but agrees well with the species described by Boisduval as Pieris
hippo (Sp. Gén. p. 534), except that it is tinged beneath with pearly ash, with scarcely any
yellow. The upper surface is sometimes rich ashy-brown, with only narrow stripes of pale
Expanse of wings varies from 2 to 2¾ inches.
6. Tachyris lycaste, Felder.
Pieris lycaste, Feld. Voy. Novara, p. 164, ♂.
Male.--Wings elongate, narrowed at the apex. Above, white, costa black-edged and dusky
towards the base; outer edge of the uppers concave, with a narrow toothed blackish border not
reaching the outer angle; lower wings with a very narrow black edging only near the outer angle.
Beneath, white, the lower wings faintly tinged with ochreous-yellow, with a narrow yellow
edging at the base; the margins as above, and a rather broad costal band pale brown.
Female.--Above, like T. hippo. Beneath, paler, lower wings pearly-ash, with a faint posterior
marginal band, and the costal and subcostal nervures light brown.
Expanse of wings ♂ 2 3/4, ♀ 2 5/8 inches.
Hab.--Celebes (Coll. Wall.).
I had described this species before I received Dr. Felder's work. He was not acquainted with
[[p. 366]] 7. Tachyris formosana, Wallace.
Pieris formosana, Wall. Proc. Zool. Soc. 1866, p. 356.
Hab.--Formosa (Coll. Wall.); Siam (Coll. W. W. Saunders).
This species has the lower wings of the male beneath edged and washed with yellow,
showing a beautiful transition from the preceding species to T. hippo and T. ada. A specimen
from Siam in Mr. W. W. Saunders' Collection agrees closely with this, and one in the British
Museum (locality unknown) has a pale sulphur-yellow tint spreading over most of the surface of
8. Tachyris andrea, Eschscholtz.
Pieris andrea, Esch. Voy. Kotzebue, pl. x. f. 23 a, b, ♂.
Female.--Above, dusky, the uppers with two white oval spots at the end of the cell as in the
Timor form, and three elongate patches separated by the median nervules; the lower wings with a
diffused discoidal white band. Beneath, as in the Timor specimens, but the posterior band is
rather wider, and the costa of the hind wings has a margin of rich yellow, which is enlarged at the
outer angle, where it joins the dark band.
Hab.--Philippine Islands (Coll. Wall., W. W. Saunders ♀).
The male figured by Eschscholtz has very elongated and pointed fore wings.
9. Tachyris hippo, Cramer.
Papilio hippo, Cr. 195 B, C, ♀. Pieris hippo, Bd. Sp. Gén. p. 534; Voll. Mon. Pier. p. 42.
Papilio phryne, Fab. Ent. Syst. III. i. p. 196, ♂. Pieris eleonora, Bd. Sp. Gén. p. 481.
Hab.--N. India, Ceylon, Singapore, Sumatra, Philippine Islands (Coll. Wall., B. M.).
This species is distinguished from its allies by the clear ochre-yellow colour of the under
surface of the lower wings in both sexes. A female from N. India in Mr. Moore's Collection has
the wings coloured nearly as in the male, except that the apical spot is absent; the under side of
the hind wings is ochre-yellow. The Ceylonese specimens are more richly coloured beneath.
10. Tachyris enarete, Boisduval.
Pieris enarete, Bd. Sp. Gén. p. 480, ♂; Guér. Voy. Favorite, pl. ii. f. 1.
Hab.--Borneo (Coll. Wall.).
Well described by Boisduval, who gives "Moluccas" as the [[p. 367]] locality on the
authority of M. Feisthamel. This is doubtless an error, as I took it in Borneo, and it is so closely
allied to the T. hippo of Malacca and Sumatra.
11. Tachyris scyllaria, MacLeay.
Pieris scyllaria, MacLeay, King's Australia, p. 459; Bd. Sp. Gén. p. 482.
I have not seen a specimen of this species.
12. Tachyris ada, Cramer.
Papilio ada, Cr. 363 C, D, ♂. Pieris ada, Bd. Sp. Gén. p. 479; Voll. Mon. Pier. p. 41, pl. 5, f.
3, ♀. Pieris cilla, Feld. Voy. Novara, p. 165.
Hab.--Bouru, Ceram, Amboina, Mysol, Aru Islands, New Guinea (Coll. Wall.).
This fine species scarcely differs in all the above localities where I myself captured it.
Boisduval describes the male well, but errs as to the female, which he had probably never seen.
This is well described and figured by Vollenhoven.
Dr. Felder describes the Aru form as a distinct species, and it certainly presents more marked
differences than those of the other localities; some specimens from Bouru and Ceram approach it
very closely, but it is generally larger in size, and in both sexes has much more and deeper
coloured rich orange tint in the abdominal region beneath; there is also a pale or nearly white
spot near the outer angle; yet these differences seem hardly definite enough to characterize a
13. Tachyris clavis, n. sp.
Male.--Differs from T. ada only by the rather broader posterior band of the hind wings both
on the upper and under surfaces.
Female.--Above, differs from T. ada in the ground colour being white, not at all tinged with
yellow or green, but rather bluish, and with a violet shade across the base of the upper and the
middle of the under wings, and in the borders being much broader, of a browner colour, and less
dentated. Beneath, there are similar differences, the chief being the much larger posterior border
on the hind wings, and the basal half being entirely yellow, with [[p. 368]] orange patches near
the anal and outer angles. Size the same as T. ada.
Hab.--Ké Island (Coll. Wall.).
The female of this form differs so strikingly in appearance from that of T. ada, which is
nearly constant through a wide range of localities, that I am induced to separate it, although, like
the various forms allied to T. hippo, the differences are not such as are usually considered to be
14. Tachyris abnormis, n. sp. (Pl. VIII. fig. 5, ♀.)
Female.--Fore wings with the outer margin more rounded than in the female of T. ada.
Above, pearly white, the body and adjacent parts of the wings with a faint greenish tinge; upper
wings with the costal margin dusky, and a large subtriangular apical patch extending from the
middle of the costa to the outer angle blackish-brown; lower wings with a border of the same
colour, widest at the outer angle and not reaching to the anal angle. Beneath, upper wings as
above, with the base ochre-yellow, and a small elongate patch of cinnabar-red, situate half way
down the cell close to the subcostal nervure; under wings entirely brown-black, the base tinged
with ochreous-yellow, which extends a short distance along the costal and anal margins.
Expanse of wings 2¾ inches.
Hab.--New Guinea (N. W. extremity) (Coll. Wall.).
This very remarkable species appears to come best here. In colouration it bears a striking
general resemblance to the beautiful nymphalideous butterfly, Mynes Geoffroyi, which inhabits
the same country.
15. Tachyris panda, Godart.
Pieris panda, Godt. Enc. Méth. ix. p. 147; Bd. Sp. Gén. p. 485; Voll. Mon. Pier. p. 44. Pieris
sulphurea, Voll. Mon. Pier. p. 32, pl. 4, f. 4, ♀.
Hab.--Java, ♂, ♀, B. M. (Horsfield).
This species appears to be confined to the island of Java. It is well described by Godart and
Boisduval, and there can be no doubt that Vollenhoven's sulphurea is the female, and the
locality, "Moluccas," which he gives from Reinwardt, an error. It is represented in the other
western islands and in Celebes, by T. nathalia, Feld., which also occurs in Java, most likely
inhabiting a separate district in that island.
[[p. 369]] 16. Tachyris nathalia, Felder.
Pieris nathalia, Feld. Wien. Ent. Monats. vi. p. 285.
Hab.--Java, Singapore (♂, ♀), Sumatra, Borneo, Celebes (♂), Philippine Islands (♂) (Coll.
Male.--Above, creamy white with a faint greenish tinge; upper wings with the costal and
outer margins narrowly black-bordered, and the apex somewhat dusky. Beneath, bright
ochre-yellow; all but the apex and basal part of the uppers much paler or whitish, costal and
outer margins finely black-edged. Head and neck olivaceous, thorax and abdomen ashy. The
specimens from Celebes are paler and have a much narrower dark border. Those from the
Philippines have a rather broader border, and a narrow one on the hind wings.
Expanse of wings 2 1/2 inches.
Female.--Above, sulphur-yellow; upper wings with the basal third dusky; a black border
from near the base of the uppers to the anal angle of the lower wings broader on the uppers and
enlarged at the apex, its outline toothed within near the outer angle of the lowers. Beneath, upper
wings with a black-curved band from the middle of the costa to the outer angle, the disc
sulphur-yellow, the base deep yellow, the apex light violet-ash with a yellow margin; lower
wings pale ochre-yellow, with a faint violet tinge, with a narrow border of rich yellow from the
base to the anal angle.
Expanse of wings 2 3/8 inches.
A specimen from Singapore and one from Java exactly agree with this description. It also
exactly agrees with Vollenhoven's figure and description of sulphurea (T. panda, ♀), except in
possessing the narrow border of bright yellow beneath. Some Singapore specimens are very pale
yellow above, and have the lower wings tinged with ochre-yellow beneath. Dr. Felder's
description of the male refers to a Malacca specimen; the Philippine form having a distinct
border to the hind wings, and a dusky patch near the outer angle of the fore wings beneath, is
almost sufficiently distinct, in such a group as this, to deserve a separate name.
17. Tachyris paulina, Cramer.
Papilio paulina, Cr. 110 E, F, ♀. Catophaga leis, Hübn. Zutr. 771, 772, ♀. Pieris neombo,
Bd. Sp. Gén. p. 539, ♀; Moore, Cat. Lep. E. I. C. pl. II. a, f. 3. [[p. 370]] Pieris albina, Bd. Sp.
Gén. p. 480, ♂. Pieris galene, Feld. Voy. Novara, p. 165, ♂ var. Pieris darada, Feld. ib. p. 166, ♂ var.
Hab.--(Males.) N. India, S. India, Penang, Java (Horsf., B. M.), Celebes, Bouru, Ceram,
Gilolo, Batchian (Coll. Wall.), Obi?, Morty?, Philippines (Wall.).
(Females, white.) N. India, Bengal, Ceylon, Philippines (W. W. Saund.), Bouru (Wall.), Obi,
Morty (Voll.), Timor, Lombock, Amboina (Wall.).
(Females, yellow.) Batchian, Aru Islands, Makian (Wall.), Java (Horsf.).
The males vary from pure white with only a dusky costa, to such as have a dusky outer
border extending up the nervures of the upper wings, and formed of small spots at the end of the
nervures of the hind wings; and the females show every gradation from the delicately coloured
form of the Moluccas (neombo, Bd.) to the darkest specimens of paulina, Cr. All the males,
however, have the under surface of two tints--white and pale ochre, and the females all have a
glossy tint of pearly white on the underside of the hind wings. The yellow form is described by
Boisduval as a variety of neombo. The specimen from Gilolo, described by Vollenhoven (Mon.
Pier. p. 36), seems intermediate, having the upper side yellow, while the under side is pearly
white, the disc of the uppers only being yellow. I am informed by Mr. Semper that he possesses
also from the Philippines specimens of these two forms with some which connect them.
18. Tachyris Rouxii, Boisduval.
Pieris Rouxii, Bd. Sp. Gén. p. 481, ♂.
Hab.--Punjaub, N. India (B. M., Coll. Hewitson, ♂).
This differs from T. paulina, ♂ (albina, Bd.), only in having the under surface uniform white.
19. Tachyris psyche, Felder.
Pieris psyche, Feld. Voy. Novara, p. 166.
Hab.--New Caledonia (Feld.), New Hebrides (Coll. W. W. Saunders).
This form differs from the rest by the lower wings and the apex of the uppers being "saffron
[[p. 371]] 20. Tachyris galathea, Felder.
Pieris galathea, Feld. Verh. z.-b. Ges. Wien. 1862, p. 485; Voy. Novara, p. 165.
This appears hardly different from T. paulina (T. albina and T. leis).
21. Tachyris ega, Boisduval.
Pieris ega, Bd. Sp. Gén. p. 536, ♂; Guér. Voy. Favorite, pl. ii. f. 2, ♂. P. melania, Bd. Sp.
Gén. p. 537 (nec Fab.), ♀.
Hab.--Australia (Coll. Wall., B. M.), New Caledonia (B. M.).
Specimens from Moreton Bay have the under side of the lower wings of the female richly
bordered with orange both on the upper and anal margins. A female from New Caledonia in the
British Museum is small, and on the under side has the uppers yellow at the base, and the disc of
the lower wings uniform rich ochre-yellow. Two others from Australia have the disc of the lower
wings beneath white, narrowly edged with orange. A small male from Lord Howe's Island, in the
British Museum, has the wings more acute and only one spot on the uppers. This is decidedly not
the Papilio melania of Fabricius, which has not yet been properly identified, and probably never
will be.1 It is therefore better to omit it altogether, since we most likely possess the same insect
under some other name.
22. Tachyris urania, n. sp.
Male.--Above, white; base of uppers and costa to the middle gray, thence with a black
border to near the outer angle, broader at the apex and extending in points along the radial and
median nervules; an indistinct black line connects these points across the apex, and a sublunate
spot between the second and third median nervules; lower wings entirely white. Beneath: upper
wings white, their apex and the lower wings pale lemon-yellow; the costa, the extreme apex, and
the transverse line faintly dusky; the sublunate spot as above, but connected to a smaller
indistinct spot at the extremity of the second median nervule; lower wings [[p. 372]] with a group
of dusky atoms just below the discoidal cell, and another barely visible near the outer angle.
Expanse of wings 2 9/10 in.
Hab.--Tondano, mountains of N. Celebes (Coll. Wall.).
This fine species has the hind wings broad and elongated almost as in T. paulina, but it is
most nearly allied to T. Jacquinotii.
23. Tachyris Jacquinotii, Lucas.
Pieris Jacquinotii, Luc. Rev. et Mag. de Zool. 1852, p. 326, ♂; Voll. Mon. Pier. p. 43. P. zoe,
Voll. Mon. Pier. p. 37, pl. 4, f. 5, ♂. P. agave, Feld. Wien. Ent. Monats. vi. p. 286, ♂, ♀.
Female.--Above, very like T. paulina, but the base of the lower wings as well as the uppers
is dusky, and the black border is broad at the outer angle, and diminishes to a point at the anal
angle, being bounded above by a bluish shade in reversed proportions, which indents it between
the nervures. Beneath, differs from T. paulina chiefly by the hind wings being pale ochre-yellow,
with a posterior submarginal angulated band not reaching the anal angle, and the space beyond
this is dusky-yellow.
Expanse of wings 2¼ - 2½ inches.
Hab.--(Males.) Batchian, Bouru, Ceram, Philippines (Coll. Wall.); New Caledonia (Coll. W.
W. Saunders). (Females.) Batchian, Ceram (Wall.); Philippines (Feld.).
I have specimens connecting T. Jacquinotii and T. zoe, and the females from the two
localities do not differ. The Philippine male is rather more yellow beneath, and the New
Caledonian still deeper yellow, but even the Moluccan specimens differ considerably in tint.
24. Tachyris alope, Wallace.
Pieris amasene, Bd. Sp. Gén. p. 535, ♂ (nec Cr.).
Female.--Differs from that of T. Jacquinotii above only by having the costa more broadly
black, and by the marginal band on the hind wings being broader and not bounded above by a
bluish shade. Beneath differs only by the transverse apical band being broader.
Hab.--(Males.) Java, Sumatra, Borneo (Coll. Wall.); India. (Females.) Java (Wall.); Sumatra
and India (B. M.); India (Moore), uniform yellow on lower wings beneath.
[[p. 373]] Vollenhoven (Mon. Pier. p. 35) describes as the female of this, that which I
consider a variety of paulina, and which Boisduval described as neombo. It is true these resemble
each other more than the pairs I have placed together, but that proves little. Horsfield paired them
as I do, and the analogy of the sexes of T. ega, from Australia, is in its favour.
The Indian males and one from Sumatra (B. M.) are much deeper yellow beneath, but the
markings agree exactly.
25. Tachyris amarella, n. sp. (Pl. IX. fig. 2, ♀.)
Female.--Wings rounded, outer margin of uppers concave. Above, sulphur-yellow; costa
and base of uppers dusky, a broad black border occupying the outer half of both the wings,
angulated on the inner margins; a row of five yellowish spots across the uppers, the second and
third largest. Beneath: the dark band is violet-brown, paler near the margin, the spots
violet-white, and there are two more indistinct spots near the outer angle; the base of the upper
wings is orange, and the disc of the lower whitish, covered with light-brown scales.
Expanse of wings 2 1/10 inches.
Hab.--New Caledonia (one specimen, B. M.).
Probably near T. alope.
26. Tachyris acrisa, Boisduval.
Pieris acrisa, Bd. Bull. Soc. Ent. Fr. 1859, p. 156, ♂.
"Size and form of P. amasene, (Bd., nec Cramer), which it completely resembles above.
Beneath, it is entirely different from all the species of this group; the whole underside of the hind
wings and the tip of the fore wings are a clear yellow-ochre, with a row of submarginal black
Hab.--Woodlark Island (East of N. Guinea).
Differs from T. alope as T. psyche does from T. paulina.
27. Tachyris leptis, Felder.
b. Liberia group.
Pieris leptis, Feld. Voy. Novara, p. 163. P. paulina, Bd. Sp. Gén. p. 538 (nec Cramer).
Hab.--Java, Borneo, N. India (♂, ♀, B. M., Coll. Wall.).
My specimen from Borneo is rather larger than the type from Java, and has a narrow black
border only near the outer angle of the lower wings. Boisduval's description of both sexes is
good, but [[p. 374]] he omits to mention that the underside of the female is slightly dusky, caused
by a few groups of ashy scales, and that the yellow margin is very indistinct. This is decidedly
not the paulina of Cramer, whose figure represents a different insect, to which Boisduval's
description of the female perhaps refers. I had named and described this species before receiving
Dr. Felder's work.
28. Tachyris celestina, Boisduval. (Pl. VIII. fig. 6, 6 a, ♀.)
Pieris celestina, Bd. Sp. Gén. p. 484; Hew. Ex. Butt. vol. ii. Pieris IV. f. 29, 30, ♂; Voll.
Mon. Pier. p. 46.
Female.--Above, bluish-white or rich yellow; basal triangle dusky; broad border and apical
triangle black, with four subapical spots of the colour of the disc. Beneath, the markings as
above; in the white variety the disc pearly-white, the base of the uppers and the abdominal
margin of the hind wings yellow, the costal margin of the hind wings chrome yellow, the apex of
the upper and the outer margin of the hind wings purplish-ashy, the white spots diffused; in the
yellow variety the border is deep and nearly uniform purplish-brown, the disc and spots of the
uppers yellow as above, the disc of the hind wings pale ochre, with the costal and abdominal
margins yellow. Head and thorax blackish, with dusky hairs. Antennæ dusky, obscurely ringed
with white beneath.
Hab.--(Males.) New Guinea, Salwatty, Waigiou, Mysol, Aru Islands (Coll. Wall.).
(Females.) Aru Islands, Waigiou (Coll. Wall.).
The specimens described by Boisduval and Vollenhoven as females are males, as can be
easily seen by examining the anal valves; specimens exactly similar are in my collection. From
the close resemblance of the male to T. Jacquinotii, except in colour, I am strongly inclined to
believe that the specimens above described are females of this species, a view which the
localities above given renders probable. I obtained several of these females of both colours,
varying somewhat in details, but evidently forming two distinct types of that sex, as occurs in T.
paulina, and in both sexes in T. cycinna.
29. Tachyris clementina, Felder.
Pieris clementina, Feld. Sitz. Akad. Wiss. xl. p. 448, ♂; Voy. Novara, tab. xxv. f. 6.
Hab.--Amboina (Coll. Felder).
[[p. 375]] Female.--Above, exactly like T. Jacquinotii, ♀, but the black posterior border is
not regularly scalloped between the nervures. Beneath, upper wings marked as in T. celestina ♀,
but tinged with greenish-yellow, the tip deep ochre-yellow; lower wings ochre-yellow, the band
and obscure submarginal spots (sometimes obsolete) as in T. celestina ♀.
Hab.--Ceram (Coll. Wall. and W. W. Saunders).
If I have rightly paired the sexes of T. celestina, then this is no doubt the female of P.
clementina, Amboina and Ceram having in almost every case the same species of insects.
30. Tachyris athama, Lucas. (Pl. IX. fig. 1, ♀.)
Pieris athama, Luc. Rev. & Mag. de Zool. 1852, p. 336, ♀; Voy. au Pôl. Sud, Lep. pl. 1, f.
10, 11; Voll. Mon. Pier. p. 36.
Female.--Upper wings pointed, outer margin concave. Above, deep sulphur-yellow,
sometimes orange-tinged; costal margin dusky, a broad jet-black marginal band forming a large
triangle at the apex of the uppers, its inner border notched and enclosing three yellow spots, two
close together near the apex, and one near the middle of the margin; band of the lower wings
angulated within, leaving a narrow toothed yellow edging near the outer angle, and terminating at
the anal angle. Beneath: disc of upper wings yellow, their base orange, their tip violet, with three
whitish spots at the edge of the transverse notched black band; lower wings pearly-white, with a
broad marginal band, violet without and nearly black within, a small spot on the outer and anal
angles, and an edging along the inner margin rich chrome-yellow.
Expanse of wings 2¼ - 2 3/8 inches.
Hab.--New Caledonia (B. M., Coll. Hew.); "Balaou, N. Guinea" (Jacquinot).
A very beautiful insect, near the female of T. celestina. Lucas describes it as "blanc jaunâtre"
above, and it is so represented in the "Voy. au Pôl. Sud," which also gives the hind border too
narrow, so that the insect is hardly recognizable.
31. Tachyris cynisca, n. sp.
Female.--Wings triangular, somewhat acute. Above, white, with a blackish border from the
middle of the costa to the anal angle, of the same form on the uppers as in T. Jacquinotii ♀, base
of uppers dusky-grey, deepening along the costa into the border, two very indistinct pale spots
near the apex. Beneath: uppers [[p. 376]] white, with the dark band browner, the apex
ashy-violet, base of the wings yellow tinged; lower wings pearly-white, the border glossy
purplish-ash, uniform; the inner margin with a fine yellow edging.
Expanse of wings 2 1/8 inches.
Hab.--Bouru (Coll. Wall.).
This is probably the female of a species allied to some of the preceding. It combines the
characters of several of them.
32. Tachyris eumelis, Boisduval.
Pieris eumelis, Bd. Sp. Gén. p. 516.
Hab.--New Ireland (Voy. Coquille).
Appears to be near T. cynisca. I have not seen a specimen.
33. Tachyris panthea, n. sp.
Form and size of T. nathalia ♀.
Female.--Above, sulphur-yellow, the base broadly dusky-black, the costa and outer margin
to the anal angle broadly black.
Beneath, like T. nathalia ♀, but the disk of the upper wings yellow, the apical band much
wider; the hind wings with a broader pearly border, and a submarginal dusky sinuated band not
reaching the outer or the anal angles.
Hab.--Philippine Islands (Coll. Hewitson and B. M.).
The yellow colour is very powdery and easily rubs off, so that worn specimens are white.
The male will be like either T. nathalia or T. zamboanga.
34. Tachyris cycinna, Hewitson.
Pieris cycinna, Hew. Ex. Butt. vol. ii. Pieris IV. f. 23, 26, ♂. P. ocina, Hew. l. c. f. 24, 25, ♀.
Hab.--Aru Islands (Coll. Wall.).
In my collection I have two males and two females, forming two pairs, which agree exactly,
except that on the underside the lower wings and the base of the uppers are in one pair pure
yellow, in the other bright orange. As they are all from the same locality it is probably one of
those cases in which a decided variation occurs in the same brood, with no intermediate links.
[[p. 377]] 35. Tachyris corinna, n. sp.
Male.--Above, exactly like T. liberia. Beneath, the upper wings have a black border along
the outer margin, broadest at the apex and regularly narrowing to the outer angle; just within this
at the apex is a spot yellower than the rest of the wing; the lower wings are uniform
chrome-yellow, with a narrow black outer border, whereas there is a mere edging in T. liberia,
which has a border of light yellow.
Expanse of wings 1 3/8 inch.
Female.--White, base of wings broadly dusky, shading into a broad black border, which is
broadest at the apex and reaches the anal angle. Beneath: the border is of the same width and
purplish, the disk of the uppers is greenish-white, with the base yellow, the lower wings are deep
Expanse of wings 1 1/8 inch.
Hab.--Waigiou (Coll. Wall.).
While the male so closely resembles T. liberia that it seems to have been considered the
same species, the female differs altogether, being more like T. cycinna (ocina, Hew.).
36. Tachyris liberia, Cramer.
Papilio liberia, Cr. 210 G, H, ♂. Pieris liberia, Godt. Enc. Méth. ix. p. 814; Bd. Sp. Gén. p.
484; (nec Voll.).
Female.--Above, bluish-ashy, the basal half and costa dusky; upper wings with a rather
narrow black border, broadest at the apex and narrowing to the outer angle, its inner edge
scalloped between the nervures; lower wings with a very narrow outer border, slightly produced
along the nervures. Beneath, upper wings greenish, their tips and the whole surface of the lower
wings chrome-yellow; two or three dusky spots near the outer angle at the margin; a very fine
black edging round the wings. Hind margin straighter than in P. eliada ♀.
Expanse of wings 2¼ inches.
Hab.--Amboina (Coll. Wall.).
This species, well figured by Cramer, appears to be restricted to the small island of Amboina,
since upon the adjacent island of Ceram another species is found which extends to Batchian, and
which Vollenhoven supposed to be T. liberia, owing to his not having an Amboina specimen
with which to compare it. The [[p. 378]] rich chrome-yellow of the under surface in both sexes at
once distinguishes Cramer's species.
37. Tachyris eliada, Hewitson.
Pieris eliada, Hew. Ex. Butt. vol. ii. Pieris IV. f. 27, 28, ♂. P. liberia, Voll. Mon. Pier. p. 45,
pl. 5, f. 4, ♀ ( nec Cram.).
Hab.--Batchian, Ceram (Wall.), Obi (Voll.)
38. Tachyris placidia, Stoll.
Papilio placidia, Stoll, Suppl. Cr. pl. 28, f. 4, 4c, ♂. Pieris placidia, Bd. Sp. Gén. p. 483;
Voll. Mon. Pier. p. 46, pl. 5, f. 5, ♀.
Hab.--Amboina, Batchian, Ceram, Morty, Waigiou (Wall.); Gilolo, Ternate (Voll.).
A curious species, with a peculiarity of colour hardly to be found elsewhere among
39. Tachyris fatima, Vollenhoven.
c. Nero group.
Pieris fatima, Voll. Tijd. v. Ent. 1866, p. 59, pl. 2, f. 1, 2, ♀.
Hab.--Celebes (Leyden Mus.).
This fine new species seems closely allied to the hitherto isolated T. placidia.
40. Tachyris nero, Fabricius.
Papilio nero, Fab. Ent. Syst. III. i. p. 153. Pieris nero, Bd. Sp. Gén. p. 485. Pieris thyria,
Horsf. Zool. Journ. iv. p. 69, t. 4, f. 2.
Hab.--N. India, Singapore, Sumatra, Java, Borneo (Coll. Wall.).
The specimens vary considerably in tint and in size, and slightly in form, but there are no
well marked races in any of the above localities. The Borneo specimens seem to have rather
acuter and more falcate wings.
41. Tachyris galba, n. sp.
Male.--Wings much more acute than T. nero, but less so than [[p. 379]] T. zarinda; outer
margin of fore wings concave to the apex; anal angle slightly produced. Above, colour the same
as T. nero; upper wings with the base and costa dusky; a black border rather broad at the apex
and vanishing at the outer angle, extending in points on the nervures, which are all black; a row
of fine dusky sagittate spots on the nervures within the border, and a small angular spot at the
upper corner of the discoidal cell. Lower wings with a narrow interrupted border, formed of spots
across the ends of the nervures, from which a black line extends a short distance along each; base
of the wings dusky. Beneath, hardly differs from T. nero, except that the submarginal spots on
the upper wings are less defined and further from the margin.
Expanse of wings 3¼ inches.
Hab.--N. India (Coll. Moore).
This fine insect has a very distinct form from either of its allies, and probably inhabits some
limited region of N. India.
42. Tachyris domitia, Felder.
Pieris domitia, Feld. Wien. Ent. Monats. vi. p. 285.
Hab.--Luzon, Philippine Islands (Coll Wall.).
Hardly distinguishable from T. nero. Has rather broader fore wings, and generally a distinct
dusky band across the hind wings beneath; the first and second median nervules of the upper
wing are not black, as they seem always to be in T. nero.
43. Tachyris zarinda, Boisduval.
Pieris zarinda, Bd. Sp. Gén. p. 486, pl. 18, f. 4.
Hab.--Celebes, Sula Islands (Coll. Wall.).
This fine species varies much in colour, like T. nero.
44. Tachyris bouruensis, n. sp.
Female.--Above, red; base of all the wings dusky black, paler on the lower wings
downwards to the anal angle, extending to the end of the cell in the uppers, where it sends off a
prolongation at right angles to near the border, and then branches downwards in a waved
submarginal line, distinct on the upper but almost obsolete on the lower wings; the costa
narrowly black to the apex, which is black, whence a border of moderate width, and dentate on
its inner edge, extends to the anal angle. Beneath: upper [[p. 380]] wings light red, their apex and
the lower wings pale reddish-ochre; a transverse irregular band beyond the end of the cell
blackish, the outer border and a submarginal band on the lower wings dusky.
Wings rather elongated, their expanse 3 1/8 inches.
Hab.--Bouru (Coll. Wall.).
This insect may not improbably be the female of T. zarinda, some species of birds being
common to Celebes and Bouru; but as it is quite as likely to be a distinct species I have thought it
better to describe and name it. The specimen is unique in my collection.
45. Tachyris zamboanga, Felder.
Pieris zamboanga, Feld. Wien. Ent. Monats. vi. p. 285, (♂ ?); Voy. Novara, tab. xxiv. f. 2, 3.
"Near melania, Fab.," Feld. The figure in the Novara Voyage shows the female form of anal
extremity, and it seems to me to be allied to T. celestina ♀. If, however, it be, as Dr. Felder says,
a male, then I cannot help thinking that T. asterope is the female of it, and that it serves to
connect the celestina group with T. nephele.
46. Tachyris asterope, Felder.
Pieris asterope, Feld. Wien. Ent. Monats. vi. p. 286, ♀.
Hab.--Luzon (Coll. Wall.).
"Near T. zamboanga; above, like T. margarita, Hübn."(Feld.). Very like T. celestina ♀, to
which it may be allied.
47. Tachyris ithome, Felder.
Pieris ithome, Feld. Wien. Ent. Monats. iii. p. 4, tab. 3, f. 1, ♂; Voll. Mon. Pier. p. 39, pl. 5,
f. 1, ♀.
Hab.--Macassar, S. Celebes (Wall.); N. Celebes (Voll.).
This species and the next form a little group of very peculiarly marked Pieridæ.
48. Tachyris nephele, Hewitson.
[[p. 381]] d. Pandione group.
Pieris nephele, Hew. Ex. Butt. vol. ii. Pieris V. f. 33, ♂; Feld. Wien. Ent. Monats. v. p. 299,
vi. p. 287, ♀.
Hab.--Luzon, Philippine Islands (Coll. Wall.).
Allied to T. ithome, which both sexes resemble.
49. Tachyris pandione, Hübner.
Hipposcritia pandione, Hübn. Zutr. 651, 652, ♂. Pieris pandione, Bd. Sp. Gén. p. 537, ♀.
Pieris paulina, var., Voll, Mon. Pier. p. 33.
Hab.--Java (B. M., Coll. Wall.).
50. Tachyris Lucasii, Wallace.
Pieris pandione ♂, Bd. Sp. Gén. p. 537 (nec Hübner). P. paulina, var., Voll. Mon. Pier. p. 33.
Male.--Above, almost exactly the same as T. leptis, Feld. (paulina, Bd.) and T. indra,
Moore, ♂; the apex of the wings is, however, more produced, and the outer margin slightly
concave, the hind margin and the ends of the nervures are finely dusky. Beneath, the black band
across the uppers is shaped nearly as in T. leptis, but it has a sagittate white spot within it at the
lower part; there is a faint dusky patch at the base of the cell, and the apex is amber-brown; the
lower wings are closely speckled with dusky yellow-brown, across which are several darker
waved or zigzag bands. The spot on the disco-cellular nervule is whitish and indistinct.
Expanse of wings 2 6/10 inches.
Hab.--Java (Coll. Wall.).
This species is very distinct from that with which it has been confounded. It is much nearer
the male of T. indra, but that species differs slightly in form, and has a black disco-cellular spot
on the underside of the lower wings.
51. Tachyris indra, Moore.
Pieris indra, Moore, Proc. Zool. Soc. 1857, p. 103, pl. 44, f. 5, ♀.
Male.--Above very like T. leptis, but the band on the uppers is rather narrower and does not
pass the first median nervule, the white spots vary from two to four; the hind wings have only a
trace of black at the outer angle, and a minute black marginal dot, sometimes obsolete, between
each pair of nervules. Beneath, the upper wings are white, greenish at the base, and pale
ochre-brown at the apex, with the transverse black band much narrower than in T. leptis, the
lower wings are pale ochre-yellow, [[p. 382]] with groups of dusky atoms disposed in irregular
macular bands, and a black disco-cellular spot.
Expanse of wings 1 3/8 inch.
Hab.--N. India (Coll. Wall., B. M.).
Pieris colimba, Bd., quoted by Moore, appears to be a MS. name, and should be altogether
52. Tachyris phbe, Felder.
Pieris phbe, Feld. Wien. Ent. Monats. v. p. 299; Voy. Novara, tab. xxv. f. 5, ♂.
Hab.--Luzon, Philippine Islands (B. M.).
This species comes nearest to T. indra.
53. Tachyris zamora, Felder.
Pieris zamora, Feld. Wien. Ent. Monats. vi. p. 286, ♂.
Hab.--Mindoro, Philippine Islands (Coll. Semper).
Dr. Felder considers this only a local modification of phbe.
54. Tachyris lalage, Doubleday.
e. Polisma group.
Pieris lalage, Doubl. Gen. of Diur. Lep. pl. 6, f. 3, ♀. P. durvasa, Moore, Proc. Zool. Soc.
1857, p. 103, pl. 44, f. 6 (err. 4), ♂.
Hab.--N. India (Coll. Moore, B. M.).
The finest species of this group; common in collections from Darjeeling.
55. Tachyris polisma, Hewitson.
Pieris polisma, Hew. Ex. Butt. vol. ii. Pieris VI. f. 38, ♂; vol. iii. Pieris VIII. f. 55, ♀; Voll.
Tijds. v. Ent. 1866, pl. 1, f. 3, 4, ♀.
Hab.--Macassar (Wall.); N. Celebes. (Voll.).
This curious little butterfly differs from every other species of Pieridæ in the shortness of its
discoidal cells. It agrees, however, with T. pandione and allies, in having the tuft of bristles at the
base of the anal valves of the male very well developed.
56. Tachyris illana, Felder.
PRIONERIS, n. g.
Pieris illana, Feld. Wien. Ent. Monats. vi. p. 287, ♂. P. ægis, Feld. lib. cit. v. p. 299; Voy.
Novara, pl. xxiv. f. 1, ♀.
Hab.--Mindanao, Philippines (Coll. Wall.).
[[p. 383]] The occurrence of a species closely allied to T. polisma in Mindanao, is another
example of the zoological connexion of this island and Celebes. T. illana marvellously resembles
Pieris cynis, although it differs in important structural characters, such as the length of the
discoidal cell, the structure of the anal valves, and the form of the club of the antennæ. I have
little doubt but that P. ægis of Felder is the female of this species.
Antennæ long, with a gradually thickened obconic club; head small, forehead very hairy,
with the blunt terminal joint of the palpi not projecting much beyond it; body stout, anal valves
of the male large, ovate, not tufted.
Upper wings triangular, the costa thickened and strongly toothed throughout its entire length;
first and second subcostal nervules arising near together at some distance before the end of the
cell, the second reaching the apex of the wing, the third of considerable length, and forming a
wide angle with the nervure; the upper radial as a branch of the subcostal, the upper
disco-cellular springing obliquely from the subcostal as if part of the second radial, and giving a
peculiar arched form to the cell of the upper wings. Cell of the hind wings two-thirds the length
of the wing.
This genus comprises a few large and fine Indian species, which seem to have no direct
affinity with the rest of the old genus Pieris. They are well characterized by the extraordinary
serrated costa, which in the larger species may be seen by the naked eye. The arrangement of the
nervures of the upper wing is also very peculiar, and I am informed that Mr. Watson had already
noticed the absence of the characteristic plumules from the wings of the males of this group,
although they exist in all the other species of Pieris.
The species in several cases seem to mimic those of the genus Thyca. P. thestylis ♂, on the
underside, most strikingly imitates T. belladonna, while the female resembles it on the upper side
as well; P. sita is a perfect representation of T. eucharis; P. cornelia is equally like T.
singhapura; and in all these cases the species of Thyca are very abundant, and are weak,
slow-flying insects, while the mimicking species of Prioneris are rare, and in all cases the pairs
which resemble each other inhabit the same district, and very often are known to come from the
[[p. 384]] 1. Prioneris thestylis, Doubleday.
Pieris thestylis, Doubl. Zool. Misc. p. 76.
Hab.--North India, Darjeeling (Coll. Wall., B. M.).
Female.--Almost exactly like P. seta, but rather larger, blacker, and the colour beneath a
rather greenish gamboge-yellow, without any tint of orange as in that species (Coll. Moore).
2. Prioneris seta, Moore.
Pieris seta, Moore, Proc. Zool. Soc. 1857, p. 102, t. 44, f. 3, ♀.
Male.--Above, white; upper wings with the costa, apex, and outer margin nearly to the outer
angle, black, and all the veins beyond the cell black-margined; lower wings with a black fringe,
and the orange markings of the under surface showing through. Beneath, the costal border is
wider and with the apex is of a pale violet slate-colour, enclosing a white spot in the cell, and one
or two others near the apex; the lower wings have patches of rich orange or chrome-yellow in the
cell and between all the nervures, except two slaty patches, one below and the other above the
base of the cell, the upper one enclosing a yellow spot.
Expanse of wings 2½ - 2¾ inches.
This appears to be the most common form, but a series of variations occur by which it is
brought nearer and nearer to P. thestylis.
In the extreme form in this direction, the upper wings above are almost exactly as in P.
thestylis; the lower wings differ in having only a very narrow dusky border, and in being of a
different tint, owing to the orange colour of the under surface showing through. Beneath, the
markings are almost identical in the two species, except that in P. seta the spots are all larger,
especially the first from the abdominal margin, which extends down to and encloses the marginal
spot, and in being all pale chrome or orange instead of pure gamboge-yellow, and the ground
colour being of a purplish slate-colour, instead of nearly black.
Expanse 3¼ inches. (P. thestylis 3 5/8 inches).
Female.--The type is probably that represented by Mr. Moore. Mr. Watson, of Manchester,
has, however, sent me for examination a curious specimen, which I believe to be a female
approximating to the colouration of the male. Above, the upper wings differ from the male in
having several dusky lines in the cell, leaving two or three distinct white spots near its upper
border; the hind wings have a narrow black toothed border, and between the anal angle and the
cell a large roundish ill-defined orange [[p. 385]] patch. Beneath, the colouring is nearly as in the
larger male specimens. The body is wanting, but the smooth costa shows it to be a female.
Expanse 3 inches.
Hab.--N. India, Darjeeling, Moulmein (Coll. Moore, Wall., B. M.).
A very puzzling species, but in most specimens quite distinct from P. thestylis, and always
smaller and of a different colour. If the extreme form only remained (and it seems now most
numerous) it would not be thought very closely allied to the species from which we can now
hardly separate it.
3. Prioneris sita, Felder.
Pieris sita, Feld. Voy. Novara, p. 161, tab. xxv. f. 12.
Wings broad, the uppers triangular, with the outer margin straight.
Male.--Above, white; the upper wings with the veins black and gradually thickened towards
the outer margin, where they spread and unite in a border; lower wings with the veins black and
thickened near the outer and hind margin, only dilated at the ends and connected by a narrow
black edge; there is a very faint dusky wavy line parallel to the outer margin on both wings.
Beneath, as above, but the veins black and thickened on both wings, the apex of the uppers and
the whole disc of the lower wings bright yellow, and round the hind margin of the latter a row of
six light cinnabar-red spots, separated from the disc by a narrow black border, the spots narrowly
bordered and notched with white.
Expanse of wings 3¼ inches.
Hab.--Ceylon (B. M., Coll. Wall. & Moore).
This remarkable species so closely resembles in marking and colour the common Thyca
eucharis of India, more especially on the under surface, that it might easily be mistaken for it. I
had described it as P. eucharoides before receiving Dr. Felder's work, and fear that Mr. Moore's
name of seta for an allied species will create some confusion.
4. Prioneris clemanthe, Doubleday.
Pieris clemanthe, Doubl. Ann. Nat. Hist. xvii. p. 23. P. Helferi, Feld. Voy. Novara, p. 161,
tab. xxv. f. 10, 11.
Hab.--North India (Coll. Saunders and Wall.).
Dr. Felder's figure seems to represent only a slight modification of the ordinary form of this
[[p. 386]] Female.--Above: upper wings dusky, with whitish bands in the cell and parallel to
the inner margin, and two curved rows of linear whitish spots beyond, the inner ones larger, more
ovate, and better marked; lower wings creamy-white, with a broad dusky outer border, which is
prolonged along the nervures to the cell, and encloses a row of faint whitish rounded spots.
Beneath, upper wings as above; lower wings as in the male, but the yellow tint paler and
extending over the spots at the outer angle. (B. M.)
5. Prioneris berenice, Lucas.
Pieris berenice, Luc. Rev. & Mag. de Zool. 1852, p. 324.
This seems very near P. clemanthe; I give the author's description:
"Near autothisbe. Wings white, the uppers with a black border formed of united spots; hind
wings slightly yellow-tinged towards the base and abdominal margin, with a narrow black border
extending on the nervures towards the outer angle in triangular patches. Beneath, the uppers as
above, the nervures black-bordered, the hind wings bright orange-brown, with the veins and
border deep black, and a marginal row of white spots; the base marked with a red spot."
The passages italicised show marked differences from P. clemanthe. I have not seen a
6. Prioneris Vollenhovii, n. sp. (Pl. IX. fig. 3, ♂.)
Form of wings as in P. thestylis.
Male.--Above, as in P. clemanthe, but the apex rather more dusky, and the posterior border
wider, with the terminations of the veins more deeply bordered. Beneath, upper wings as above;
lower wings with the nervures all black and thickened, the basal two-thirds and four spots at the
outer angle rich yellow, the anal angle black, extending in a dusky border to the middle of the
hind margin, at the base of the wing a hairy red patch as in P. clemanthe.
Expanse of wings 3 3/8 inches.
Hab.--Borneo, Sarawak (Coll. Wall. and W. W. Saunders).
7. Prioneris cornelia, Vollenhoven.
Pieris cornelia, Voll. Mon. Pier. p. 5, pl. 2, f. 2.
Hab.--Borneo (Leyden Museum).
This species in form and colouring resembles Thyca singhapura, [[p. 387]] although it differs
so remarkably in structure. I have not seen a specimen.
8. Prioneris philonome, Boisduval.
Pieris philonome, Bd. Sp. Gén. p. 453, ♂.
Hab.--Java (Coll. Payen).
A rare species, allied to P. clemanthe and P. Vollenhovii by the curious hairy red patch at the
base of the lower wings beneath. I have not seen a specimen.
9. Prioneris autothisbe, Hübner.
Pieris autothisbe, Hübn. Samml. Ex. Schmett. ii. tab. cxxiii.; Bd. Sp. Gén. p. 452; Voll. Mon.
Pier. p. 20, pl. 3, f. 5, ♀.
Hab.--Java (Wall.). (B. M. ♀.)
I took this species at 4,000 feet elevation on the mountains of West Java. Vollenhoven has
made known the female, which by its black upper wings shows its affinity with P. thestylis.
The Eastern species of Eronia form a compact and homogeneous group of very closely allied
forms, several of which are hardly more than slight geographical modifications. These insects
frequent most generally the dry and open forests and plantations. They fly tolerably quickly, with
an undulating movement, and often settle upon flowers. The clear blue tints with which the males
are adorned render them striking objects in the forests of the Malay Islands.
1. Eronia avatar, Moore.
Eronia avatar, Moore, Cat. Lep. E. I. C. p. 61, pl. ii. a, f. 1, ♂.
Hab.--N. India, Darjeeling (Coll. Wall.).
Female.--Apex rather more rounded than in the male; dusky, with the spaces between the
nervures whitish, as in E. valeria and iobæa, but leaving only the nervures and outer margin
(enclosing the submarginal row of spots) dusky. Beneath, silvery, with the nervures obscurely
dusky, the median nervure and its branches on the upper wing broadly blackish (Coll. Moore).
This fine species seems to be an extreme development of the valeria type.
[[p. 388]] 2. Eronia valeria, Cramer.
Papilio valeria, Cr. 85 A. Pieris valeria, Bd. Sp. Gén. p. 444. Eronia valeria, Voll. Mon.
Pier. p. 56.
Hab.--Malacca, Sumatra, Java, Borneo, Lombock, Flores (Coll. Wall.).
In this species the male is nearly constant, while the female, as in the whole genus, varies
exceedingly, often more or less closely imitating species of Danais. The Bornean males have
rather more elongate fore wings, with a slightly concave outer margin.
3. Eronia hippia, Fabricius.
Papilio hippia, Fab. Ent. Syst. III. i. p. 59; Don. Ins. India, pl. 25, f. 1. Danais hippia, Godt.
Enc. Méth. ix. p. 193. Pieris valeria, var. A, Bd. Sp. Gén. p. 445. Eronia gæa, Feld. Voy.
Novara, p. 190.
Hab.--India (Coll. Wall., B. M.)
This species has been generally confounded with E. valeria, but it is quite as distinct as most
of the other species of the genus. It is undoubtedly the hippia of Fabricius and Godart. The
female differs less from the male than in any other species of the genus.
4. Eronia bbera, Eschscholtz.
Paphia bbera, Esch. Voy. Kotzebue, vol. iii. (♀ fig.). Eronia bbera, Feld. Wien. Ent.
Monats. vi. p. 288, ♂.
Hab.--Philippine Is. (Luzon and Mindoro). (Coll. Wall., B. M.)
Closely allied to E. hippia.
5. Eronia ceylanica, Felder.
Eronia ceylanica, Feld. Voy. Novara, p. 191.
Hab.--Ceylon (Coll. Wall.).
Barely distinct from E. hippia.
6. Eronia argolis, Felder.
Eronia argolis, Feld. Wien. Ent. Monats. 1860, p. 52; Voll. Mon. Pier. p. 58, pl. 7, f. 2, ♀.
Hab.--Batchian, Gilolo (Coll. Wall.).
A richly-marked species. The female, as in several others of the genus, is often more or less
tinged with yellow.
[[p. 389]] 7. Eronia tritæa, Felder.
Eronia tritæa, Feld. Wien. Ent. Monats. vi. p. 288, t. 3, f. 2, ♂; Voll. Mon. Pier. p. 58, pl. 7,
f. 3, ♀.
Hab.--Celebes (Macassar and Menado). (Coll. Wall.).
8. Eronia phocæa, Felder.
Eronia phocæa, Feld. Wien. Ent. Monats. v. p. 299, ♂; vi. p. 288, ♀.
Hab.--Mindanao (Coll. Wall.).
Near E. tritæa and E. argolis.
9. Eronia iobæa, Boisduval.
Pieris iobæa Bd. Voy. Astrol. p. 57, pl. 2, f. 5, 6, ♂ (err.♀); Sp. Gén. p. 445. Eronia iobæa,
Voll. Mon. Pier. p. 57, pl. 7, f. 1, ♀.
Hab.--Ceram, Waigiou, New Guinea (Coll. Wall.).
The males do not vary perceptibly; the females vary in colour from bluish-white to pale
yellow or ochre-yellow, and the markings from being as prominent as in the specimen figured by
Vollenhoven to others in which the whole surface is brown-black, with a submarginal row of
spots, and faint traces of the usual markings.
This African genus is restricted in the East to Southern Central and North Western India, and
Ceylon. It is the tropical form of our northern Anthocharis.
1. Callosune eucharis, Fabricius.
Papilio eucharis, Fab. Syst. Ent. p. 472 (nec Drury). Anthocharis eucharis, Bd. Sp. Gén. p.
Hab.--India, Ceylon (B. M.); Deccan (Coll. Moore).
2. Callosune danae, Fabricius.
Papilio danae, Fab. Syst. Ent. p. 476. Anthocharis danae, Bd. Sp. Gén. p. 570.
Hab.--Central India, Ceylon (B. M.).
[[p. 390]] 3. Callosune etrida, Boisduval.
Anthocharis etrida, Bd. Sp. Gén. p. 576.
Hab.--Punjaub, N. India, Ceylon (B. M.).
In the Ceylon specimen the border of the hind wings is continuous, otherwise it agrees with
the Indian form.
(Anthocharis daphalis, Moore, Proc. Zool. Soc. 1865, p. 491, pl. 31, f. 14, from Kunawur, N.
W. India (10,000 feet alt.), belongs to the Palæarctic fauna.)
This northern genus ranges over the higher parts of the Himalayan range as far as Assam, and
one species has been recently described from the Nilgherries. Of the nine species here introduced
not more than three or four can be properly said to belong to the Indian region. One is described
from the Sandwich Islands.
1. Colias Fieldii, Ménétries.
Colias Fieldii, Ménétries, Cat. Imp. Acad. p. 79, t. 1, f. 5.
Hab.--N. W. Himalayas (Coll. Moore, B. M.).
A geographical form of C. myrmidone.
2. Colias xenodice, Felder.
Colias xenodice, Feld. Wien. Ent. Monats. iii. p. 394.
Hab.--Assam (Feld.); Nepal (Wall., ex Coll. Hardwicke).
Barely distinct from C. edusa or C. myrmidone.
3. Colias eogene, Felder.
Colias eogene, Feld. Voy. Novara, p. 196, pl. xxviii. f. 7.
Hab.--Himalaya (Coll. Felder).
Near C. Boothii, Curtis.
4. Colias Ponteni, Wallengren.
Colias Ponteni, Wallengren, Wien. Ent. Monats. iv. p. 33.
"Alis supra fulvido-flavis, maris iridatis, macula discoidali anticarum, limbo latissimo
omnium, costisque extra cellulam, nigris; apice anticarum flavo-maculato, omnibus maris infra
fulvido-flavis, posticis apiceque anticarum fminæ grisescente [[p. 391]] flavis; anticis circulo
discoidali ferrugineo, posticis macula ferruginea discoidali, cosla flava intersecta margineque
interiore late nigro-pustulato." (Wallengren.)
Hab.--Honolulu, Sandwich Islands.
5. Colias hyale, Linnæus.
Papilio hyale, Lin. S. N. i. p. 469. Colias hyale, Bd. Sp. Gén. p. 650.
Hab.--N. India, Nepal (Coll. Moore, B. M.).
6. Colias neriene, Fischer.
Colias neriene, Fischer, Ent. Russ. Lep. t. 11, f. 3, 4; Bd. Sp. Gén. p. 646.
Hab.--N. W. Himalayas (6,000-10,000 feet elevation, Capt. Lang).
7. Colias nilagiriensis, Felder.
Colias nilagiriensis, Feld. Wien. Ent. Monats. iii. p. 395.
Hab.--Malabar (Nilgherries). (Coll. Felder.)
A geographical modification of C. neriene.
8. Colias shipkee, Moore.
Colias shipkee, Moore, Proc. Zool. Soc. 1865, p. 492, pl. xxxi. f. 13.
Hab.--N. W. India (13,000 to 15,000 feet alt., Capt. Lang).
9. Colias ladakensis, Felder.
Colias ladakensis, Feld. Voy. Novara, p. 197, pl. xxvii. f. 8, 9, ♂.
Hab.--Himalaya, Ladak, Spiti (Dr. Stolickza). (Coll. Felder.)
This African and desert genus is represented only by five species in North Western, Western
and Southern India.
1. Idmais calais, Cramer.
Papilio calais, Cr. 53 C, D, 351 A, D. Idmais calais, Bd. Sp. Gén. p. 587. Pieris amata,
Godt. Ent. Méth. ix. p. 131.
Hab.--Madras, S. India (B. M.).
2. Idmais phisadia, Godart.
Pieris phisadia, Godt. Enc. Méth. ix. p. 132. Idmais phisadia, Bd. Sp. Gén. p. 587.
Hab.--N. W. India (B. M.).
[[p. 392]] 3. Idmais dynamene, Klug.
Pontia dynamene, Klug, Symb. Phys. Ins. t. 7, f. 5, 6. Idmais dynamene, Bd. Sp. Gén. p. 588.
Hab.--Punjaub (B. M.).
4. Idmais fausta, Olivier.
Papilio fausta, Oliv. Voy. en Syrie, t. 33, f. 4. Idmais fausta, Bd. Sp. Gén. p. 586.
Hab.--N. W. India (B. M.)
5. Idmais fulvia, n. sp. (Pl. IX. fig. 5, ♂, ♀.)
Near I. fausta.
Male.--Upper wings with the outer margin more convex, the apex blacker, leaving a row of
small spots, the discoidal spot very large and nearly round; thorax and abdomen blackish.
Beneath, paler in colour, the markings more distinct, and the discoidal spot much larger than in I.
Female.--Resembles the male, but the brick-red colour is replaced by white.
Expanse of wings, 2 inches.
Hab.--S. India (Moore); Scinde (B. M.).
This is an especially Indian genus. In the Archipelago one species is found in Sumatra, two
in Java, and one in each of the small islands as far as Timor. It is not known from Borneo or from
any other parts of the Archipelago. It seems to inhabit only mountainous and rather dry districts,
and this has probably determined its peculiar distribution. Many of the species are very variable,
especially in the female sex, and I have had much difficulty in grouping them satisfactorily.
1. Thestias venilia, Godart.
Pieris venilia, Godt. Enc. Méth. ix. p. 121. Thestias venilia, Bd. Sp. Gén. p. 594; Voll. Mon.
Pier. p. 51. Papilio ænippe, Cr. 157 C, D, ♀ ?.
Hab.--Java (Horsf.); Timor ? (Voll.).
This distinct species is probably confined to Java, where it is very scarce.
[[p. 393]] 2. Thestias Reinhardtii, Vollenhoven.
Thestias Reinhardtii, Voll. Tijd. v. Ent. iii. p. 126; Mon. Pier. p. 50, pl. vi. f. 1, ♂.
Female.--Above, upper wings black, the internal border broadly white, the base dusky, a
white oblique spot across the origin of the subcostal nervules, a smaller one below it, and three
ovate spots near the outer margin; lower wings white, the base dusky, and a broad black outer
border. In one specimen the markings of the upper wings are more developed, and of a yellow
colour. Beneath, upper wings pale yellow, more or less irrorated with dusky, a large brown
discoidal spot, beyond which is an irregular brown band, merging in a large patch at the outer
angle; lower wings ochre-yellow, sparely irrorated on the disc, and densely on the outer margin, a
small discoidal spot, and between it and the border a row of six or seven rufous-brown spots,
dusky-margined, the third and fourth largest, and the whole nearly confluent.
Hab.--Baly, Lombock, Flores (Coll. Wall.).
A well-marked species, variable in the female, and restricted to the islands between Java and
3. Thestias balice, Boisduval.
Thestias balice, Bd. Sp. Gén. p. 593, ♂.
Hab.--Java (B. M.), ♂.
4. Thestias Vollenhovii, Wallace.
Thestias balice, Voll. Mon. Pier. p. 50, ♂, ♀ (nec Boisd.).
Hab.--Timor (Coll. Wall.), ♂, ♀.
Vollenhoven's description is correct. Boisduval's refers to the Java form, which is
5. Thestias venatrix, n. sp.
Male.--Above, yellow; the hind wings with a very narrow dark edge, extending slightly on
the nervures at the outer angle; fore wings with a large triangular orange-red patch in the middle,
surrounded by a black border, narrow within and on the costa, wider at the apex and outer
margin, the nervures and a discoidal spot black; the base of the wing bluish, and of the costa
brown. Beneath: upper wings yellow, their margins and the whole of the lower wings pale
yellow, thickly irrorated with reddish-ochre, discoidal spot on the lower wings of the same colour
and small, on the upper wings larger and black.
Smaller than T. balice, to which it is allied.
Hab.--Moulmein (B. M.).
[[p. 394]] 6. Thestias marianne, Cramer.
Papilio marianne, Cr. 217 C, D, E. Thestias marianne, Bd. Sp. Gén. p. 592, ♂, ♀.
Hab.--India (Coll. Wall., B. M.).
In a female from Madras, the transverse black band does not pass below the end of the cell,
and some of this form in the British Museum are small, and have a narrow border; in another,
from Bengal, the whole upper surface is tinged with ochre-yellow. One from the Punjaub is very
large and finely coloured.
7. Thestias pyrene, Linnæus.
Papilio pyrene, Linn. Syst. Nat. ii. p. 762. P. pyrene, Cr. 125 A, B, C, ♂. Thestias pirene, Bd.
Sp. Gén. p. 593, ♂. Papilio ænippe, Drury, Exot. Ent. pl. 5, f. 2, ♂ (good). P. ænippe, Cr. 229 B,
C. Papilio sesia, Don. Ins. China, pl. 31, f. 2 (bad).
Male.--Drury's figure is the best, and serves well to identify the species. It differs from
Cramer's pyrene above, by the wings being more acute, the orange patch less regular, and the
black border to the hind wings generally wider. Beneath, it varies from clear yellow--with a
small black discoidal spot on each wing, a blackish spot at the outer angle of the uppers, and two
or three indistinct orange spots between the discoidal spot and the margin of the lower wings--to
ochreish-yellow, thickly speckled with black on the costal region of the uppers and the whole
surface of the lower wings, and the two or three orange submarginal spots more diffused. It
differs strikingly from Cramer's pyrene in the entire absence of the row of brown spots with
Female.--Something like Cramer's ænippe (229 B, C), and perhaps the same (not 105 C, D).
It varies greatly in size and colouring, as pointed out by Mr. Moore, from Capt. Lang's notes. It
is always white and black, or nearly so, and offers the greatest difference of the sexes in this
Expanse of wings (male) 2¼ - 2¾ inches; (female) 2¼ - 2 7/8 inches.
Hab.--N. India, Bengal, China (B. M., Coll. Wall.).
The largest specimens of both sexes are from Darjeeling. Those from the N. W. Himalayas
are of moderate size, while those from the plains of Punjaub and Oude are smaller and less richly
coloured. These last are hardly distinguishable from some specimens of T. pirenassa. The two in
fact may well be con- [[p. 395]] sidered as one abundant and variable species, which has become
segregated into several forms, which may be divided into two pretty well-marked groups.
There has been great confusion about this species. Drury's is the best and most recognizable
figure I have seen. The Linnæan description is so imperfect that the name ought perhaps not to be
retained, but it has been so generally applied to some of the forms of this species that I think it
better to keep to it.
8. Thestias pirenassa, n. sp. (Pl. IX. fig. 4, ♂.)
Thestias ænippe, Bd. Sp. Gén. p. 591, ♀ (as ♂) ?, (nec Cr., Dru.).
Male.--Costa much curved, hind wings subtriangular.
Above, like T. pyrene, but the transverse black band always touches the discoidal spot of the
uppers. Beneath, yellow, faintly speckled with black; on the hind wings a row of six or seven
spots, of which one on the inner margin, as well as the third and fourth, are larger and have
pearly or violet-white ocelli.
Female.--Very like the male, differing only in the orange patch being smaller and less
regular, and having two black spots on its lower part. Beneath, as in the male, but the spots are
more distinct, and almost all have white ocelli.
Expanse of wings (♂ and ♀), 2 inches.
Hab.--Bombay, Madras (Coll. Wall., Moore).
Variety A.--Is rather clearer in colour, has the upper wings rather more acute, and a trace
only of border on the hind wings in both sexes. Beneath, of a richer chrome-yellow tint. The
female generally as in the type, but sometimes almost white as in the extreme form of T. pyrene
Expanse of wings, ♂ 2 inches; ♀ 1 3/4- 2 inches.
Hab.--Sylhet, Darjeeling (Coll. Wall., Moore).
Variety B. Male.--Hind wings rather less triangular, transverse band completely enclosing
the discoidal spot, under surface clear yellow, with minute discoidal spots, as in the extreme
form of T. pyrene ♂. Female.--Pale yellow, with a broader dusky border, the spot on the upper
wings smaller and more divided by the nervures, either the same yellow as the disc or more or
less tinged with orange; beneath, ochreish or chrome-yellow, marked as in the female of T.
Expanse of wings, ♂ 2 inches; ♀ 2 - 2 1/4 inches.
Hab.--Bengal, N. India (Coll. Moore).
[[p. 396]] 9. Thestias Ludekingii, Vollenhoven.
Thestias Ludekingii, Voll. Tijd. v. Ent. iii. p. 125; Mon. Pier. p. 49, pl. 5, f. 6, ♀.
Hab.--Sumatra (Leyden Museum).
A distinct species, allied to T. pirenassa.
This genus consists of the largest and most gaudily-coloured Pieridæ, and is especially
characteristic of the Malayan Islands. For more than eighty years it had consisted of two species
only, when I had the good fortune to discover two more in the Moluccas, and the group now
comprises six sufficiently distinct species, all of which are found in the Malay Archipelago, one
of them only ranging out of it over all tropical Asia. The Moluccas and Timor are the eastern
limits of the group, no specimen having yet been found in New Guinea.
1. Iphias glaucippe, Linnæus.
Papilio glaucippe, Lin. S. N. ii. p. 762; Cr. 164 A, B, C. Iphias glaucippe, Bd. Sp. Gén. p.
Hab.--(Type).--India, Sumatra, Celebes (Coll. Wall., B. M.). (Var.).--Java, Flores, Timor,
Philippine Is. (Coll. Wall., B. M.).
This species has a wide range and varies slightly in each locality in form, size, and
colouration. These variations may be best grouped under two forms,--the type, which has the
orange patch bordered with black on the inner side,--the variety which has little or no black
separating the orange from the white of the wings. The Celebes specimens are very large and
have very acutely-pointed wings. The Celebes form has most black, the Timor form least; and if
the species became extinct over the remainder of its range, these two would undoubtedly be
considered distinct, since they differ constantly both in form and markings.
2. Iphias borneensis, n. sp.
Iphias glaucippe, var., Wall. Journ Ent. vol. 2, p. 3.
Male.--Like I. glaucippe, but the orange apical patch much reduced, leaving a wide border
all round it, which at the apex encroaches on the black spots; lower wings sometimes with a
submarginal row of small spots.
Female.--Like I. glaucippe, but the apical patch small and [[p. 397]] white, the disc of the
hind wings only tinged with sulphur-yellow, and the margin more broadly black.
Hab.--Borneo (Coll. Wall.), (♀ B. M.).
This interesting species bears the same relation to glaucippe that leucogynia does to
leucippe; the female being coloured white and black only.
3. Iphias Vossii, Maitland.
Iphias Vossii, Mait. Tijd. voor Ent. ii. p. 25; Voll. Mon. Pier. p. 55, pl. vi. f. 4.
Hab.--Island of Nias, near Sumatra (Leyden Mus.).
4. Iphias sulphurea, Wallace.
Iphias sulphurea, Wall. Journ. of Ent. vol. 2, p. 5. I. Felderi, Voll. Mon. Pier. p. 53, pl. 6, f.
2, 3, ♂, ♀.
Hab.--Batchian, Gilolo, Morty Island (Coll. Wall.).
5. Iphias leucogynia, Wallace.
Iphias leucogynia, Wall. Journ. of Ent. vol. 2, p. 4, pl. 1, f. 1 ♂, f. 2 ♀.
Hab.--Bouru (Coll. Wall.).
6. Iphias leucippe, Cramer.
Papilio leucippe, Cr. 36 A, B, C. Iphias leucippe, Bd. Sp. Gén. p. 596.
Hab.--Amboina, Ceram (Coll. Wall.).
This magnificent insect is not uncommon in Amboina as well as in Ceram, although the
Leyden Museum possesses no specimens from the former locality.
A small group of insects separated from Gonepteryx by Dr. Boisduval, and now generally
adopted. Their wings are larger in proportion to their bodies, their antennæ more slender, the
wing cells much shorter, and the radial of the lower wing is very differently situated. Dercas
gobrias flies slowly in forest clearings near the ground, often settling in damp places. The genus
seems confined to India, Borneo, and Sumatra.
[[p. 398]] 1. Dercas gobrias, Hewitson.
Gonepteryx gobrias, Hew. Trans. Ent. Soc. 3rd ser. vol. ii. p. 246, pl. xvi. f. 1, ♂; Butler,
Proc. Zool. Soc. 1865, p. 432, pl. xxv. fig. 4, ♂, ♀.
Female.--Larger than the male, pale yellow, the discoidal spot and transverse streak almost
Hab.--Sumatra, Borneo (Coll. Wall., B. M.).
2. Dercas Verhuellii, Van der Hoeven.
Colias Verhuellii, Van d. Hoeven, Tijd. Nat. Gesch. 1838, t. 8, f. 3, 4, ♂. Rhodocera lycorias,
Doubl. Zool. Misc. 1842, p. 77 (wood cut) ♀.
Hab.--N. India, Silhet (Coll. Wall., B. M.).
The female has the apex of the wings more falcate and acute than the male, and the markings
and outline of the wings vary considerably.
In the "Genera of Diurnal Lepidoptera," Mr. Doubleday puts his R. lycorias ♀ as a synonym
of D. Verhuellii, but retains lycorias ♂ as a distinct species from Silhet. As no specimen
answering to his description exists in the British Museum, or in any private collection with which
I am acquainted, I am inclined to think that the specimen was probably compounded of the fore
wings and body of D. Verhuellii, with the hind wings of a Callidryas.
3. Dercas Wallichii, Doubleday.
Gonepteryx Wallichii, Doubl. Proc. Ent. Soc. 1848, p. xlvii. G. urania, Butler, Proc. Zool.
Soc. 1865, p. 458, pl. xxvi. f. 5.
Hab.--N. India (B. M.).
This species agrees better with Dercas than with Gonepteryx, by the short discoidal cells,
though it is somewhat intermediate between the two genera.
Of this genus two species of the European type enter the Indian region on its north-western
boundary, and these are probably confined to the higher ranges of the Himalayas. A third species
has been described by Dr. Felder, from Northern China.
[[p. 399]] 1. Gonepteryx nipalensis, Doubleday.
Gonepteryx nipalensis, Doubl. Gen. Diurn. Lep. p. 71. G. rhamni, G. R. Gray, Lep. Ins.
Nepaul. t. 5, f. 1 (nec Linn.).
Hab.--N. India, Darjeeling (B. M.).
2. Gonepteryx zaneka, Moore.
Gonepteryx zaneka, Moore, Proc. Zool. Soc. 1865, p. 493, pl. xxxi. f. 18.
Hab.--N. W. Himalayas (B. M.).
"In thick forests about Narkunda, 50 miles from Simla." (Capt. Lang).
3. Gonepteryx acuminata, Felder.
Gonepteryx acuminata, Feld. Wien. Ent. Monats. vi. p. 23.
Hab.--Ning-po, China (Coll. Felder).
A variety of G. aspasia, Men., from the Amur.
This wide-spread tropical genus is fairly represented in the Malay Islands. Most of the
species are very abundant in individuals, frequenting the open country, gardens and river banks,
and in the dry season settling by hundreds together on wet and muddy places, or on the extreme
edge of the sinking waters of the streams. Some of the named species are distinguished by such
slight characters that it is possible they may in some cases be different forms (perhaps different
broods) of the same species.
1. Callidryas thisorella, Boisduval.
Callidryas thisorella, Bd. Sp. Gén. p. 609.
Hab.--India, Australia (Queensland). (Coll. Wall., B. M.)
A female from Australia only differs from the male above in the larger discoidal spot.
Beneath, it is of a pinkish pearly tint, with the irrorations obsolete, a large dusky discoidal spot
and another small one on each wing, and a faint submarginal line formed of dusky patches on
2. Callidryas philippina, Cramer.
Papilio philippina, Cr. 361 C, D. Callidryas philippina, Bd. Sp. Gén. p. 619.
Hab.--Bengal, Central India. (Coll. Wall., B. M.)
[[p. 400]] 3. Callidryas pyranthe, Linnæus.
Papilio pyranthe, Lin. S. N. ii. p. 763. Callidryas pyranthe, Bd. Sp. Gén. p. 611. Papilio
alcyone, Cr. 58 A, B, C.
Hab.--N. India, Malacca, Sumatra, Java, Flores, China, Formosa, Philippine Islands (Coll.
The specimens from Formosa are very large, but agree in colour and marking with those
from other localities; another specimen from the same place is as remarkably small.
4. Callidryas minna, Boisduval.
Callidryas minna, Bd. Sp. Gén. p. 612.
Hab.--India, Flores (Wall.); Australia (Voy. Novara).
I took a specimen of what seems to be this species in Flores. It has a less acute apex and a
more rounded outer margin of the fore wings than in C. pyranthe, and the border is more
regularly curved within, and extends farther along the costa.
5. Callidryas alcmeone, Cramer.
Papilio alcmeone, Cr. 141 E. Callidryas alcmeone, Bd. Sp. Gén. p. 627. ? Papilio catilla, Cr.
229 D, E.
Hab.--India, Malacca, Sumatra, Java, Borneo, Philippine Islands, Celebes, Moluccas, Timor
6. Callidryas hilaria, Cramer.
Papilio hilaria, Cr. 339 A, B. Callidryas hilaria, Bd. Sp. Gén. p. 626. Papilio jugurtha, Cr.
187 E, F.
Hab--India, Malacca, Borneo, Java, Lombock, Timor, Batchian, Australia (Coll. Wall.).
7. Callidryas crocale, Cramer.
Papilio crocale, Cr. 55 C, D, ♀. Callidryas crocale, Bd. Sp. Gén. p. 625. Callidryas endeer,
Bd. Voy. Astrol. p. 63, pl. 2, f. 3, 4, ♀.
Male.--Above, sulphur-yellow, the outer margins paler, and the base of a different texture as
in C. alcmeone and C. hilaria, but with no marked difference of colour. Beneath, exactly like C.
alcmeone, but differs in tint as above.
Female.--Varies much, from a yellow form very like C. hilaria ♀, [[p. 401]] to the dark
insect figured by Boisduval. Beneath, the colour is in some specimens nearly uniform yellow or
buff, in others there is a distinct submarginal zigzag dusky band.
Hab.--Celebes, Philippine Islands, Batchian, Ceram, Timor, Australia, Queensland (Coll.
This has been confounded by Vollenhoven with C. alcmeone, from which, however, it is
quite distinct. The Timor specimens are very small in both sexes.
8. Callidryas phlegeus, n. sp.
Male.--Above, exactly like C. crocale ♂. Beneath, like C. hilaria ♂. Thus differing from C.
crocale just as C. hilaria differs from C. alcmeone.
Hab.--Timor (Coll. Wall.).
If C. hilaria and C. alcmeone are maintained as distinct species, we cannot refuse the same
rank to this form.
9. Callidryas gorgophone, Boisduval.
Callidryas gorgophone, Bd. Sp. Gén. p. 632.
Hab.--Australia, Queensland (Coll. Wall.).
10. Callidryas scylla, Linnæus.
Papilio scylla, Lin. S. N. ii. p. 763; Cr. 12 C, D. Callidryas scylla, Bd. Sp. Gén. p. 631.
Hab.--Java, Sumatra, Lombock, Timor, Celebes, Ternate, Batchian (Coll. Wall.).
ADDENDA ET CORRIGENDA.
Page 307, lines 2 and 13, for 35 read 37; and in the Table read as follows:--
| Groups of Islands
Page 315. In the List of New Species, dele Thestias dissimilis.
Page 316, line 23, dele "Pieris athama, Lucas = Tachyris celestina, Boisd. ♀."
Page 317. Pontia Crookera. Not in British Collections.
Page 317. Pontia medusa. I can find no record of authentic specimens.
Page 327. Terias celebensis. Expanse of wings 1 1/8 - 1 7/8 inch (not 1 7/8 inch to 2½ inches).--A. R. W.,
Note Appearing in the Original Work
1. Fabricius says, "alis posticis subtus obscure glaucis," and Godart, "le dessous des ailes inférieures
est d'un vert-blanchâtre obscur." This is therefore evidently not the P. melania of Boisduval (the ♀ of his P.
ega), which has the lower wings beneath pale yellow and rich orange. [[on p. 371]]